Origin of Species Variorum

Comparison with 1869

CHAPTER VIII.

HYBRIDISM.

Distinction between the sterility of first crosses and of hybrids — Sterility various in degree, not universal, affected by close interbreeding, removed by domestication — Laws governing the sterility of hybrids — Sterility not a special endowment, but incidental on other differences not accumulated by natural selection — Causes of the sterility of first crosses and of hybrids — Parallelism between the effects of changed conditions of life and of crossing — Dimorphism and trimorphism — Fertility of varieties when crossed and of their mongrel offspring not universal — Hybrids and mongrels compared independently of their fertility — Summary.

THE view generally entertained by naturalists is that species, when intercrossed, have been specially endowed with ... sterility, in order to prevent their confusion. .. .. .. .. This view certainly seems at first highly probable, for species within the same country could hardly have been kept distinct had they been capable of .. freely crossing.

Text in this page (from paragraph 100, sentence 210 to paragraph 100, sentence 500, word 25) is not present in 1869

CHAPTER VIII.

HYBRIDISM.

Distinction between the sterility of first crosses and of hybrids—Sterility various in degree, not universal, affected by close inter-breeding, removed by domestication—Laws governing the sterility of hybrids—Sterility not a special endowment, but incidental on other differences, not accumulated by natural selection—Causes of the sterility of first crosses and of hybrids—Parallelism between the effects of changed conditions of life and of crossing—Dimorphism and trimorphism—Fertility of varieties when crossed and of their mongrel offspring not universal—Hybrids and mongrels compared independently of their fertility—Summary.

THE view generally entertained by naturalists is that species, when intercrossed, have been specially endowed with the quality of sterility, in order to prevent the confusion of all organic forms. This view certainly seems at first probable, for species within the same country could hardly have kept distinct had they been capable of crossing freely. ↑ The importance of the fact that first crosses between distinct species and hybrids are very generally sterile, has, I think, been much underrated by some late writers. On the theory of natural selection the case is especially important, inasmuch as this sterility can hardly have been increased by the continued preservation of successive, profitable degrees of sterility. But to this subject I shall have to return, and I hope ultimately to show that this sterility is neither a specially acquired nor endowed quality, but is incidental on other acquired and little-known differences in the reproductive systems of the parent-species.