Origin of Species Variorum

Comparison with 1872

the genera within a family, be represented by a vertical line of varying thickness, crossing the successive geological formations in which the species are found, the line will sometimes falsely appear to begin at its lower end, not in a sharp point, but abruptly; it then gradually thickens upwards, sometimes keeping of equal thickness for a space, and ultimately thins out in the upper beds, marking the decrease and final extinction of the species. This gradual increase in number of the species of a group is strictly conformable with the theory; as the species of the same genus, and the genera of the same family, can increase only slowly and progressively; for the process of modification and the production of a number of allied forms must be slow and gradual,— one species giving rise first to two or three varieties, these being slowly converted into species, which in their turn produce by equally slow steps other species, and so on, like the branching of a great tree from a single stem, till the group becomes large.

On Extinction .

We have as yet spoken only incidentally of the disappearance of species and of groups of species. On the theory of natural selection, the extinction of old forms and the production of new and improved forms are intimately connected together. The old notion of all the inhabitants of the earth having been swept away at successive periods by catastrophes, is very generally given up, even by those geologists, as Elie de Beaumont, Murchison, Barrande, &c., whose general views would naturally lead them to this conclusion. On the contrary, we have every reason to believe, from the study of the tertiary formations, that species and groups of

the genera of a family, be represented by a vertical line of varying thickness, ascending through the successive geological formations in which the species are found, the line will sometimes falsely appear to begin at its lower end, not in a sharp point, but abruptly; it then gradually thickens upwards, often keeping for a space of equal thickness, and ultimately thins out in the upper beds, marking the decrease and final extinction of the species. This gradual increase in the number of the species of a group is strictly conformable with my theory, for the species of the same genus, and the genera of the same family, can increase only slowly and progressively; .. the process of modification and the production of a number of allied forms necessarily being a slow and gradual process,— one species first giving rise to two or three varieties, these being slowly converted into species, which in their turn produce by equally slow steps other varieties and species, and so on, like the branching of a great tree from a single stem, till the group becomes large.

On Extinction.

We have as yet spoken only incidentally of the disappearance of species and of groups of species. On the theory of natural selection the extinction of old forms and the production of new and improved forms are intimately connected together. The old notion of all the inhabitants of the earth having been swept away by catastrophes at successive periods is very generally given up, even by those geologists, as Elie de Beaumont, Murchison, Barrande, &c., whose general views would naturally lead them to this conclusion. On the contrary, we have every reason to believe, from the study of the tertiary formations, that species and groups of