Origin of Species Variorum

Comparison with 1872

other causes more or less local and temporary, but depend on general laws which govern the whole animal kingdom." M. Barrande has made forcible remarks to precisely the same effect. It is, indeed, quite futile to look to changes of currents, climate, or other physical conditions, as the cause of these great mutations in the forms of life throughout the world, under the most different climates. We must, as Barrande has remarked, look to some special law. We shall see this more clearly when we treat of the present distribution of organic beings, and find how slight is the relation between the physical conditions of various countries and the nature of their inhabitants.

This great fact of the parallel succession of the forms of life throughout the world, is explicable on the theory of natural selection. New species are formed by having some advantage over older forms; and the forms, which are already dominant, or have some advantage over the other forms in their own country, give birth to the greatest number of new varieties or incipient species; for these latter must be victorious in a still higher degree in order to be preserved and to survive. We have distinct evidence on this head, in the plants which are dominant, that is, which are commonest in their own homes, and are most widely diffused, producing the greatest number of new varieties. It is also natural that the domi- nant, varying, and far-spreading species, which have already invaded to a certain extent the territories of other species, should be those which would have the best chance of spreading still further, and of giving rise in new countries to other new varieties and species. The process of diffusion would often be very slow, .. depending on climatal and geographical changes, .. on strange accidents, and on the gradual acclimatisation of new species to the various climates through which they might have to pass, but in the long run the dominant forms would generally succeed in spreading and would ultimately prevail.

This great fact of the parallel succession of the forms of life throughout the world, is explicable on the theory of natural selection. New species are formed by new varieties arising, which have some advantage over older forms; and those forms, which are already dominant, or have some advantage over the other forms in their own country, would naturally oftenest give rise to new varieties or incipient species. .. .. .. .. .. .. .. .. .. .. .. .. .. .. .. .. .. .. .. We have distinct evidence on this head, in the plants which are dominant, that is, which are commonest and most widely diffused, compared with other less dominant plants, producing the greatest number of new varieties. It is also natural that the dominant, varying, and far-spreading species, which already have invaded to a certain extent the territories of other species, should be those which would have the best chance of spreading still further, and of giving rise in new countries to new varieties and species. The process of diffusion may often be very slow, being dependent on climatal and geographical changes, or on strange accidents, or on the gradual acclimatisation of new species to the various climates through which they have to pass, but in the course of time the dominant forms will generally succeed in spreading.