fact of | fact of 1859 1860 1861 1866 |  fact of 1869 1872 |  
  
fossil remains from closely consecutive 
 formations, | formations, 1859 1860 1861 1866 |  | formations 1869 1872 |  
  
 though ranked as distinct species, being closely related, | though ranked as distinct species, being closely related, 1859 1860 1861 1866 |  
| being closely related, though ranked as distinct species, 1869 1872 |  
  
is obvious.  As the accumulation of each formation has often been interrupted, and as long blank intervals have intervened between successive formations, we ought not to expect to find, as I attempted to show in the last chapter, in any one or 
 in any two | in any two 1866 1869 1872 |  
| two 1859 1860 1861 |  
  
 formations | formations 1859 1860 1861 1866 1869 |  | formations, 1872 |  
  
all the intermediate varieties between the species which appeared at the commencement and close of these 
 periods; | periods; 1859 1860 1861 1866 |  | periods: 1869 1872 |  
  
but we ought to find after intervals, very long as measured by years, but only moderately long as measured geologically, closely allied forms, or, as they have been called by some authors, representative species; and these 
 assuredly we | assuredly we 1866 1869 1872 |  | we assuredly 1859 1860 1861 |  
  
do find.  We find, in short, such evidence of the slow and scarcely sensible 
 mutation | mutation 1859 1860 1861 1866 1869 |  | mutations 1872 |  
  
of specific forms, as we have 
 a just | a just 1859 1860 1861 1866 |  | the 1869 1872 |  
  
right to 
 expect. | expect. 1866 1869 1872 |  | expect 1859 1860 1861 |  
  
 ..| ..... 1866 1869 1872 |  | to 1859 1860 1861 |  
  
 ..| ..... 1866 1869 1872 |  | find. 1859 1860 1861 |  
  
 | 
 
On
 | 
On
 1866 1872 |  | 
On
 1859 1860 1861 1869 |  
  
 
the
 | 
the
 1866 1872 |  | 
the
 1859 1860 1861 1869 |  
  
 
State
 | 
State
 1866 1872 |  | 
state
 1859 1860 |  | 
State
 1861 1869 |  
  
 
of
 | 
of
 1866 1872 |  | 
of
 1859 1860 1861 1869 |  
  
 
Development
 | 
Development
 1866 1872 |  | 
Development
 1859 1860 1861 1869 |  
  
 
of
 | 
of
 1866 1872 |  | 
of
 1859 1860 1861 1869 |  
  
 
Ancient
 | 
Ancient
 1866 1872 |  | 
Ancient
 1859 1860 1861 1869 |  
  
 
compared with Living Forms.  | 
compared with Living Forms.  1866 |  
| 
Forms
.—
 1859 1860 |  
| 
compared with Living Forms.
—
 1861 |  
| 
compared with Living Forms.
 1869 |  
| 
compared with Living Forms
.  1872 |  
  
 | 
 We have seen in the fourth chapter that the degree of differentiation and specialisation of the parts 
 in 
 all | all 1861 1866 1869 |  all 1872 |  
  
organic beings, when 
 arrived at | arrived at 1866 1869 1872 |  | come to 1861 |  
  
maturity, is the best standard, as yet suggested, of their degree of perfection or highness.  We have also seen 
 that, | that, 1866 1869 1872 |  | that 1861 |  
  
as the specialisation of parts 
 and organs | and organs 1861 1866 1869 |  and organs 1872 |  
  
is an advantage to each being, so natural selection will 
 tend | tend 1866 1869 1872 |  
| constantly tend thus 1861 |  
  
to render the organisation of each being more specialised and perfect, and in this sense higher; not but that it may 
 and will | and will 1861 1866 1869 |  and will 1872 |  
  
leave many creatures with simple and unimproved structures fitted for simple conditions of life, and in some cases will even degrade or simplify the organisation, yet leaving such degraded beings better fitted for their new walks of life. ↑| 2 blocks not present in  1861 1866 1869 1872; present in  1859 1860 |  |  There has been much discussion whether recent forms are more highly developed than ancient. 
I will not here enter on this subject, for naturalists have not as yet defined to each other's satisfaction what is meant by high and low forms. 
 |  
  
↑| 1 blocks not present in  1859 1860 1861 1866 1869 1872; present in  |  | 
 |  
   In 
 another and 
more general 
 manner, | manner, 1866 |  
| sense the more recent forms must, on my theory, be higher than the more ancient; for each 1860 |  
  
 |