Origin of Species Variorum

Comparison with 1869

exist, the species must be descended from a succession of modified varieties, which have supplanted each other, but which have never blended with other individuals or varieties of the same species; so that, at each successive stage of modification and improvement, all the individuals of the same variety will be descended from a single parent. But in the majority of cases, namely, with all organisms which habitually unite for each birth, or which occasionally intercross, ... the individuals of the same species inhabiting the same area will be kept nearly uniform by intercrossing; so that many individuals will go on simultaneously changing, and the whole amount of modification ... at each stage will not be due to descent from a single parent. To illustrate what I mean: our English racehorses differ slightly from the horses of every other breed; but they do not owe their difference and superiority to descent from any single pair, but to continued care in the selecting and training of many individuals during many generations.

Before discussing the three classes of facts, which I have selected as presenting the greatest amount of difficulty on the theory of "single centres of creation," I must say a few words on the means of dispersal.

Means of Dispersal .—

Sir C. Lyell and other authors have ably treated this subject. I can give here only the briefest abstract of the more important facts. Change of climate must have had a powerful influence on migration; an impassable region when its climate was different from what it now is, may have been a high road for migration; ... I shall, however, presently have to discuss this branch of the subject in some detail. Changes of level in the land must also have been highly influential: a narrow isthmus

exist), each species, on my theory, must have descended from a succession of improved varieties, which can never have blended with other individuals or varieties, but will have supplanted each other; so that, at each successive stage of modification and improvement, all the individuals of each variety will have descended from a single parent. But in the great majority of cases, namely, with all organisms which habitually unite for each birth, or which often intercross, I believe that during the slow process of modification the individuals of the species will have been kept nearly uniform by intercrossing; so that many individuals will have gone on simultaneously changing, and the whole amount of modification will not have been due, at each stage, to descent from a single parent. To illustrate what I mean: our English race-horses differ .. from the horses of every other breed; but they do not own their superiority and difference to descent from any single pair, but to continued care in selecting and training many individuals during each generation.

Means of Dispersal .

Sir C. Lyell and other authors have ably treated this subject. I can give here only the briefest abstract of the more important facts. Change of climate must have had a powerful influence on migration: a region when its climate was different may have been a high road for migration, but now be impassable; I shall, however, presently have to discuss this branch of the subject in some detail. Changes of level in the land must also have been highly influential: a narrow isthmus