Origin of Species Variorum

Comparison with 1869

something definite about the means of distribution, we shall be enabled to speculate with security on the former extension of the land. But I do not believe that it will ever be proved that within the recent period most of our continents which now stand quite separate, have been continuously, or almost continuously, united with each other, and with the many existing oceanic islands. Several facts in distribution,— such as the great difference in the marine faunas on the opposite sides of almost every continent,— the close relation of the tertiary inhabitants of several lands and even seas to their present inhabitants,— a certain degree of relation (as we shall hereafter see) between the distribution of mammals and the depth of the sea,— these and other such facts seem to me opposed to the admission of such prodigious geographical revolutions within the recent period, as are necessary on the view advanced by Forbes and admitted by his many followers. The nature and relative proportions of the inhabitants of oceanic islands likewise seem to me opposed to the belief of their former continuity with continents. Nor does their almost universally volcanic composition favour the admission that they are the wrecks of sunken continents;— if they had originally existed as continental mountain-ranges, some at least of the islands would have been formed, like other mountain-summits, of granite, metamorphic schists, old fossiliferous and other .. rocks, instead of consisting of mere piles of volcanic matter.

I must now say a few words on what are called accidental means, but which more properly might be called occasional means of distribution. I shall here confine myself to plants. In botanical works, this or that plant is stated to be ill adapted for wide dissemination; but

something definite about the means of distribution, we shall be enabled to speculate with security on the former extension of the land. But I do not believe that it will ever be proved that within the recent period continents which are now quite separate, have been continuously, or almost continuously, united with each other, and with the many existing oceanic islands. Several facts in distribution,— such as the great difference in the marine faunas on the opposite sides of almost every continent,— the close relation of the tertiary inhabitants of several lands and even seas to their present inhabitants,— the degree of affinity between the mammals inhabiting islands with those of the nearest continent, being in part determined (as we shall hereafter see) by the depth of the intervening ocean,— these and other such facts seem to me opposed to the admission of such prodigious geographical revolutions within the recent period, as are necessitated on the view advanced by Forbes and admitted by his many followers. The nature and relative proportions of the inhabitants of oceanic islands likewise seem to me opposed to the belief of their former continuity with continents. Nor does the almost universally volcanic composition of such islands favour the admission that they are the wrecks of sunken continents;— if they had originally existed as ... mountain-summits, of granite, metamorphic schists, old fossiliferous or other such rocks, instead of consisting of mere piles of volcanic matter.

I must now say a few words on what are called accidental means, but which more properly should be called occasional means of distribution. I shall here confine myself to plants. In botanical works, this or that plant is stated to be ill adapted for wide dissemination; but