Origin of Species Variorum

Comparison with 1861

As all the organic beings, extinct and recent, which have ever lived on this earth have to be classed together, and as all have been connected by the finest gradations, the best, or indeed, if our collections were nearly perfect, the only possible arrangement, would be genealogical. Descent being on my view the hidden bond of connexion which naturalists have been seeking under the term of the natural system. On this view we can understand how it is that, in the eyes of most naturalists, the structure of the embryo is even more important for classification than that of the adult. For the embryo is the animal in its less modified state; and in so far it reveals the structure of its progenitor. In two groups of animal, however much they may at present differ from each other in structure and habits, if they pass through the same or similar embryonic stages, we may feel assured that they have both descended from the same or nearly similar parents, and are therefore in that degree closely related. Thus, community in embryonic structure reveals community of descent. ↑ It will reveal this community of descent, however much the structure of the adult may have been modified and obscured; we have seen, for instance, that cirripedes can at once be recognised by their larvæ as belonging to the great class of crustaceans. As the embryonic state of each species and group of species partially shows us the structure of their less modified ancient progenitors, we can clearly see why ancient and extinct forms of life should resemble the embryos of their descendants,— our existing species. Agassiz believes this to be a law of nature; but I am bound to confess that I only hope

As all the organic beings, extinct and recent, which have ever lived on this earth, have to be classed together, and as all have been connected by fine gradations, the best, or, if our collections were nearly perfect, the only possible arrangement, would be genealogical; descent being on this view the hidden bond of connexion which naturalists have been seeking under the term of the natural system. On this view we can understand how it is that, in the eyes of most naturalists, the structure of the embryo is even more important for classification than that of the adult. ↑ In two or more groups of animals, however much they may at present differ from each other in structure and habits, if they pass through closely similar embryonic stages, we may feel almost assured that they have descended from the same parent-form, and are therefore ... closely related. Thus, community in embryonic structure reveals community of descent; but dissimilarity in embryonic development does not prove discommunity of descent, for in one of two groups all the developmental stages may have been suppressed, or may have been so greatly modified as no longer to be recognised, through adaptations, during the earlier periods of growth, to new habits of life. Community of descent will, however, often be revealed, although the structure of the adult may have been greatly modified and thus obscured; we have seen, for instance, that cirripedes, though externally so like shellfish, can at once be recognised by their larvæ as belonging to the great class of crustaceans. As the embryonic state of each species and group of species shows us more or less completely the structure of their less modified ancient progenitors, we can .. see why ancient and extinct forms of life should resemble the embryos of our existing species, their descendants. Agassiz believes this to be a law of nature; but I am bound to confess that I only hope