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4 blocks not present in 1859 1860 1861 1866; present in 1869 1872
An animal may possess various parts in a perfect state, and yet they may in one sense be rudimentary, for they are useless: thus the tadpole of the common Salamander or newt, as Mr. G. H. Lewes remarks, "has gills, and passes its existence "in the water; but the Salamandra atra, which lives "high up among the mountains, brings forth its young "full-formed. This animal never lives in the water. "Yet if we open a gravid female, we find tadpoles "inside her with exquisitely feathered gills; and when "placed in water they swim about like the tadpoles of the "water-newt. Obviously this aquatic organisation has "no reference to the future life of the animal, nor has "it any adaptation to its embryonic condition; it has "solely reference to ancestral adaptations, it repeats a "phase in the development of its progenitors."

pistil, which is in a rudimentary state, 1859 1860 1861 1866
rudimentary pistil, 1869 1872

with hairs as in other compositæ, for the purpose of brushing 1859 1860 1861 1866
with hairs, in the usual manner, for brushing 1869
in the usual manner with hairs, which serve to brush 1872

and conjoined anthers. 1866 1869 1872
anthers. 1859 1860 1861

developed, if of use, should not be called rudimentary: they 1861 1866
developed, if of use, should not be called rudimentary; they cannot properly be said to be in an atrophied condition; they 1860
developed, if of use, should not be considered as rudimentary: they 1869
they 1872

called nascent, and may hereafter be developed by natural selection to any further extent. 1861 1866 1869
called nascent, and may hereafter be developed to any extent by natural selection. 1860
developed, unless we have reason to suppose that they were formerly more highly developed, ought not to be considered as rudimentary. They may be in a nascent condition, and in progress towards further development. 1872

other hand, are essentially useless, as teeth which never cut through the gums. 1861 1866 1869
individuals of the same species are very liable to vary in degree of development and in other respects. Moreover, in closely allied species, the degree to which the same organ has been rendered rudimentary occasionally differs much. 1859
other hand, are essentially useless, as teeth which never cut through the gums; in a still less developed condition, they would be of still less use. 1860
other hand, are either quite useless, such as teeth which never cut through the gums, or almost useless, such as the wings of an ostrich, which serve merely as sails. 1872

1 blocks not present in 1859 1860 1866 1869 1872; present in 1861
Moreover, in closely allied species, the degree to which the same organ has been rendered rudimentary occasionally differs much.

they would be of still less use, when in a 1866
they would be of even less use, when in a 1869
organs in this condition would formerly, when 1872

condition, they cannot 1866 1869
OMIT 1872

under the present state of things have 1866
have 1869 1872

formed 1866 1869
of even less use than at present, they cannot formerly have been produced 1872

by 1866
through variation and 1869 1872

as we shall see, by 1866
by the power of 1869

is 1860 1861 1866 1869
is, however, often 1872

know what organs are nascent; 1861 1866 1869
know what are nascent organs; 1860
distinguish between rudimentary and nascent organs; 1872

looking to the future, we cannot of course tell how any part will be developed, and whether it is now in a nascent condition; looking to the past, creatures with an organ in a nascent 1866
looking to the future, we cannot of course tell how any part will be developed, and whether it is now nascent; looking to the past, creatures with an organ in a nascent 1860 1861
looking to the future, we can- not of course tell how any part will be developed, and whether it is now in a nascent condition; looking to the past, creatures with an organ in this 1869
for we can judge only by analogy whether a part is capable of further development, in which case alone it deserves to be called nascent. Organs in this 1872

pistils
pistil
are
are.
essentially
....
alike
....
in
....
nature.
....
An
organ,
organ
serving for two purposes, may become rudimentary or utterly aborted for one, even the more important
purpose;
purpose,
and remain perfectly efficient for the other. Thus in plants, the office of the pistil is to allow the pollen-tubes to reach the ovules
within
protected in
the
ovarium.
ovarium
at
at
its
its
base.
base.
The pistil consists of a stigma supported on
the
a
its
style; but in some
Compositæ,
compositæ,
the male florets, which of course cannot be fecundated, have a pistil, which is in a rudimentary state, for it is not crowned with a stigma; but the style remains well
developed
developed,
and is clothed with hairs as in other compositæ, for the purpose of brushing the pollen out of the surrounding and conjoined anthers. Again, an organ may become rudimentary for its proper purpose, and be used for a distinct
one:
object:
in certain
fishes
fish
the
swimbladder
swim bladder
swim-bladder
seems to be
rudimentary
nearly rudimentary
for its proper function of giving buoyancy, but has become converted into a nascent breathing organ or lung.
Many
Other
similar instances could be given.
Useful organs,
Organs,
however little developed, if of use, should not be called rudimentary: they may be called nascent, and may hereafter be developed by natural selection to any further extent. Rudimentary
organs
organs,
in
on
the other hand, are essentially useless, as teeth which never cut through the gums. As they would be of still less use, when in a still less
developed,
developed
condition, they cannot under the present state of things have been formed by natural selection, which
latter
....
acts solely by the preservation of useful modifications. They relate to a former
state
condition
of
things,
their possessor,
and have been
partially
....
retained
retained,
as we shall see, by inheritance. It is difficult to know what organs are nascent; looking to the future, we cannot of course tell how any part will be developed, and whether it is now in a nascent condition; looking to the past, creatures with an organ in a nascent