Origin of Species Variorum

Comparison with 1869

of a class, nothing is more common, or more necessary, in order fully to understand the relations of the parts, than the discovery of rudiments. This is well shown in the drawings given by Owen of the bones of the leg of the horse, ox, and rhinoceros.

It is an important fact that rudimentary organs, such as teeth in the upper jaws of whales and ruminants, can often be detected in the embryo, but afterwards wholly disappear. It is also, I believe, a universal rule, that a rudimentary part .. is of greater size relatively to the adjoining parts in the embryo, than in the adult; so that the organ at this early age is less rudimentary, or even cannot be said to be in any degree rudimentary. Hence .. rudimentary organs in the adult, are often said to have retained their embryonic condition.

I have now given the leading facts with respect to rudimentary organs. In reflecting on them, every one must be struck with astonishment: for the same reasoning power which tells us .. that most parts and organs are exquisitely adapted for certain purposes, tells us with equal plainness that these rudimentary or atrophied organs, are imperfect and useless. In works on natural history, rudimentary organs are generally said to have been created "for the sake of symmetry," or in order "to complete the scheme of nature." .. .. .. .. .. .. .. .. .. .. .. .. .. But this is not an explanation, merely a re-statement of the fact. Nor is it consistent with itself: thus the boa-constrictor has rudiments of hind-limbs and of a pelvis, and if it be said that these bones have been retained "to complete the scheme of nature," why, as Professor Weismann asks, have they not been retained by other snakes, which do not possess even a vestige of these same bones? What would be thought of an astronomer, who maintained that the satellites revolve in elliptic courses round their planets "for the sake of symmetry," because the planets thus revolve round the sun? An eminent physiologist accounts for the presence of rudimentary organs, by supposing that they serve to excrete matter in excess, or matter injurious to the system; but can we suppose that the

of the same class, nothing is more common, or more necessary, than the use and discovery of rudiments. This is well shown in the drawings given by Owen of the bones of the leg of the horse, ox, and rhinoceros.

It is an important fact that rudimentary organs, such as teeth in the upper jaws of whales and ruminants, can often be detected in the embryo, but afterwards wholly disappear. It is also, I believe, a universal rule, that a rudimentary part or organ is of greater size relatively to the adjoining parts in the embryo, than in the adult; so that the organ at this early age is less rudimentary, or even cannot be said to be in any degree rudimentary. Hence, also, a rudimentary organ in the adult is often said to have retained its embryonic condition.

I have now given the leading facts with respect to rudimentary organs. In reflecting on them, every one must be struck with astonishment: for the same reasoning power which tells us plainly that most parts and organs are exquisitely adapted for certain purposes, tells us with equal plainness that these rudimentary or atrophied organs are imperfect and useless. In works on natural history rudimentary organs are generally said to have been created "for the sake of symmetry," or in order "to complete the scheme of nature;" but this seems to me no explanation, merely an imposing re-statement of the fact. ↑ Would it be thought sufficient to say that because planets revolve in elliptic courses round the sun, satellites follow the same course round their planets, for the sake of symmetry, and to complete the scheme of nature? An eminent physiologist accounts for the presence of rudimentary organs, by supposing that they serve to excrete matter in excess, or injurious to the system; but can be suppose that the