Origin of Species Variorum

Comparison with 1872

as, or even sometimes more useful than, parts of high physiological importance. Rudimentary organs may be compared with the letters in a word, still retained in the spelling, but become useless in the pronunciation, but which serve as a clue .. for its derivation. On the view of descent with modification, we may conclude that the existence of organs in a rudimentary, imperfect, and useless condition, or quite aborted, far from presenting a strange difficulty, as they assuredly do on the old doctrine of creation, might even have been anticipated in accordance with the views here explained.

Summary .

In this chapter I have attempted to show, that the arrangement of all organic beings throughout all time in groups under groups— that the nature of the relationships by which all living and extinct beings are united by complex, radiating, and circuitous lines of affinities into a few grand classes,— the rules followed and the difficulties encountered by naturalists in their classifications,— the value set upon characters, if constant and prevalent, whether of high or of the most trifling importance, or, as with rudimentary organs, of no importance,— the wide opposition in value between analogical or adaptive characters, and characters of true affinity; and other such rules;— all naturally follow if we admit the common parentage of allied forms, ... together with their modification through variation and natural selection, with the contingencies of extinction and divergence of character. In considering this view of classification, it should be borne in mind that the element of descent has been universally used in ranking together the sexes, ages, dimorphic forms, and acknowledged varieties of the same species, however much they may differ from each other in structure. If we

as, or even sometimes more useful than, parts of high physiological importance. Rudimentary organs may be compared with the letters in a word, still retained in the spelling, but become useless in the pronunciation, but which serve as a clue in seeking for its derivation. On the view of descent with modification, we may conclude that the existence of organs in a rudimentary, imperfect, and useless condition, or quite aborted, far from presenting a strange difficulty, as they assuredly do on the ordinary doctrine of creation, might even have been anticipated, and can be accounted for by the laws of inheritance.

Summary .

In this chapter I have attempted to show, that the subordination of group to group in all organic beings throughout all time; that the nature of the relationship, by which all living and extinct organisms are united by complex, radiating, and circuitous lines of affinities into one grand system; the rules followed and the difficulties encountered by naturalists in their classifications; the value set upon characters, if constant and prevalent, whether of high vital importance, or of the most trifling importance, or, as in rudimentary organs, of no importance; the wide opposition in value between analogical or adaptive characters, and characters of true affinity; and other such rules;— all naturally follow on the view of the common parentage of those forms which are considered by naturalists as allied, together with their modification through natural selection, with its contingencies of extinction and divergence of character. In considering this view of classification, it should be borne in mind that the element of descent has been universally used in ranking together the sexes, ages, dimorphic states, and acknowledged varieties of the same species, however different they may be in structure. If we