Origin of Species Variorum

Why does not every collection of fossil remains afford plain evidence of the graduation and mutation of the forms of life? Although geological research has undoubtedly revealed the former existence of many links, bringing numerous forms of life much closer together, it does not yield the infinitely many fine graduations between past and present species required on my theory; and this is the most obvious and forcible of the many objections which may be urged against it. ↑ Why, again, do whole groups of allied species appear, though certainly they often falsely appear, to have come in suddenly on the several geological stages? Although we now know that organic beings appeared on this globe, at a period incalculably remote, long before the lowest bed of the Silurian system was deposited, why do we not find beneath this system great piles of strata stored with the remains of the progenitors of the Silurian fossils? For .. on my theory such strata must somewhere have been deposited at these ancient and utterly unknown epochs in the world's history.

I can answer these question and .. objections only on the supposition that the geological record is far more imperfect than most geologists believe. It cannot be objected that there has not been time sufficient for any amount of organic change; for the lapse of time has been so great as to be utterly inappreciable by the human intellect. The number of specimens in all our museums is absolutely as nothing compared with the countless generations of countless species which certainly have existed. The parent-form of any two or more species would not be in all its characters directly intermediate between its modified offspring, any more than the rock-pigeon is directly intermediate in crop and tail between its descendants the pouter and fantail pigeons. We should not be able to recognise a species as the parent