Origin of Species Variorum

Comparison with 1869

slightest advantage in certain individuals, at any age or during any season, over those with which they come into competition, or better adaptation in however slight a degree to the surrounding physical conditions, will turn the balance.

With animals having separated sexes there will be in most cases a struggle between the males for possession of the females. The most vigorous individuals, or those which have most successfully struggled with their conditions of life, will generally leave most progeny. But success will often depend on having special weapons, or means of defence, or charms; and a slight advantage will lead to victory.

As geology plainly proclaims that each land has undergone great physical changes, we might have expected to find that organic beings .. have varied under nature, in the same way as they .. have varied under ... domestication. And if there be any variability under nature, it would be an unaccountable fact if natural selection did not come into play. It has often been asserted, but the assertion is .. incapable of proof, that the amount of variation under nature is a strictly limited quantity. Man, though acting on external characters alone and often capriciously, can produce within a short period a great result by adding up mere individual differences in his domestic productions; and every one admits that species present individual differences. .. .. .. .. But, besides such differences, all naturalists admit that varieties exist, which are considered sufficiently distinct to be worthy of record in systematic works. No one has drawn any clear distinction between individual differences and slight varieties; or between more plainly marked varieties and sub-species, and species. On separate continents, and on different parts of the same continent when

slightest advantage in one being, at any age or during any season, over those with which it comes into competition, or better adaptation in however slight a degree to the surrounding physical conditions, will turn the balance.

With animals having separated sexes there will in most cases be a struggle between the males for the possession of the females. The most vigorous males, or those which have most successfully struggled with their conditions of life, will generally leave most progeny. But success will often depend on the males having special weapons or means of defence, or on their charms; and the slighest advantage will lead to victory.

As geology plainly proclaims that each land has undergone great physical changes, we might have expected that organic beings would have varied under nature, in the same way as they generally have varied under the changed conditions of domestication. And if there be any variability under nature, it would be an unaccountable fact if natural selection had not come into play. It has often been asserted, but the assertion is quite incapable of proof, that the amount of variation under nature is a strictly limited quantity. Man, though acting on external characters alone and often capriciously, can produce within a short period a great result by adding up mere individual differences in his domestic productions; and every one admits that there are at least individual differences in species under nature. But, besides such differences, all naturalists have admitted the existence of varieties, which they have considered sufficiently distinct to be worthy of record in their systematic works. No one can draw any clear distinction between individual differences and slight varieties; or between more plainly marked varieties and sub-species, and species. On separate continents, and on different parts of the same continent when