Origin of Species Variorum

Comparison with 1869

existing .. groups. Recent forms are generally looked at as being, on the whole, higher in the scale of organisation than ancient .. forms; and they must be higher, in so far as the later and more improved forms have conquered the older and less improved forms in the struggle for life; they have also generally had their organs more specialized for different functions. This fact is perfectly compatible with numerous beings still retaining .. simple and but little improved structures, fitted for simple conditions of life; it is likewise compatible with some forms having retrograded in organisation, by having become at each stage of descent better fitted for changed and degraded habits of life. Lastly, the wonderful law of the long endurance of allied forms on the same continent,— of marsupials in Australia, of edentata in America, and other such cases,— is intelligible, for generally within the same country, the existing and the extinct will naturally be allied by descent.

Looking to geographical distribution, if we admit that there has been during the long course of ages much migration from one part of the world to another, owing to former climatal and geographical changes and to the many occasional and unknown means of dispersal, then we can understand, on the theory of descent with modification, most of the great leading facts in Distribution. We can see why there should be so striking a parallelism in the distribution of organic beings throughout space, and in their geological succession throughout time; for in both cases the beings have been connected by the bond of ordinary generation, and the means of modification have been the same. We see the full meaning of the wonderful fact, which must have struck every traveller, namely, that on the same continent, under the most diverse conditions, under heat and cold, on mountain and lowland, on deserts and marshes, most

existing allied groups. Recent forms are generally looked at as being, on the whole, higher than ancient and extinct forms; and they are higher in so far as the later and more improved forms have conquered the older and less improved organic beings in the struggle for life; they will also generally have had their organs more specialised for different functions. This fact is perfectly compatible with numerous beings still retaining a simple and little improved organisation fitted for simple conditions of life; it is likewise compatible with some forms having retrograded in organisation, though becoming under each grade of descent better fitted for their changed and degraded habits of life. Lastly, the law of the long endurance of allied forms on the same continent,— of marsupials in Australia, of edentata in America, and other such cases,— is intelligible, for within a confined country, the recent and the extinct will .. be closely allied by descent.

Looking to geographical distribution, if we admit that there has been during the long course of ages much migration from one part of the world to another, owing to former climatal and geographical changes and to the many occasional and unknown means of dispersal, then we can understand, on the theory of descent with modification, most of the great leading facts in Distribution. We can see why there should be so striking a parallelism in the distribution of organic beings throughout space, and in their geological succession throughout time; for in both cases the beings have been connected by the bond of ordinary generation, and the means of modification have been the same. We see the full meaning of the wonderful fact, which has struck every traveller, namely, that on the same continent, under the most diverse conditions, under heat and cold, on mountain and lowland, on deserts and marshes, most