Origin of Species Variorum

Comparison with 1872

embryological characters are the most valuable of all. The real affinities of all organic beings, in contradistinction to their adaptive resemblances, are due to inheritance or community of descent. The Natural System is a genealogical arrangement, with the acquired grades of difference, marked by the terms, varieties, species, genera, families, &c.; and we have to discover the lines of descent by the most permanent characters whatever they may be and of however slight vital importance. .. ..

The similar framework of bones in the hand of a man, wing of a bat, fin of a porpoise, and leg of the horse,— the same number of vertebræ forming the neck of the giraffe and of the elephant,— and innumerable other such facts, at once explain themselves on the theory of descent with slow and slight successive modifications. The similarity of pattern in the wing and in the leg of a bat, though used for such different purpose,— in the jaws and legs of a crab,— in the petals, stamens, and pistils of a flower is likewise, to a large extent, intelligible on the view of the gradual modification of parts or organs, which were aboriginally alike in an early progenitor in each of these classes. On the principle of successive variations not always supervening at an early age, and being inherited at a corresponding not early period of life, we can clearly see why the embryos of mammals, birds, reptiles, and fishes should be so closely alike, and .. so unlike the adult forms. We may cease marvelling at the embryo of an air-breathing mammal or bird having branchial slits and arteries running in loops, like those of a fish which has to breathe the air dissolved in water, by the aid of well-developed branchiæ.

Disuse, aided sometimes by natural selection, will often have reduced organs when rendered useless under changed habits or .. conditions of life; and we can .. understand on this view the meaning of rudimentary organs. But disuse and selection will generally act on each creature, when it has

embryological characters are often the most valuable of all. The real affinities of all organic beings are due to inheritance or community of descent. The natural system is a genealogical arrangement, in which we have to discover the lines of descent by the most permanent characters, however slight their vital importance may be.

The framework of bones being similar in the hand of a man, wing of a bat, fin of the porpoise, and leg of the horse,— the same number of vertebræ forming the neck of the giraffe and of the elephant,— and innumerable other such facts, at once explain themselves on the theory of descent with slow and slight successive modifications. The similarity of pattern in the wing and leg of a bat, though used for such different purpose,— in the jaws and legs of a crab,— in the petals, stamens, and pistils of a flower, is likewise intelligible on the view of the gradual modification of parts or organs, which were alike in the early progenitor of each class. On the principle of successive variations not always supervening at an early age, and being inherited at a corresponding not early period of life, we .. clearly see why the embryos of mammals, birds, reptiles, and fishes are so closely similar, and are so unlike their adult forms. We may cease marvelling at the embryo of an air-breathing mammal or bird having branchial slits and arteries running in loops, like those in a fish which has to breathe the air dissolved in water by the aid of well-developed branchiæ.

Disuse, aided sometimes by natural selection, will often tend to reduce an organ, when it has become useless by changed habits or under changed conditions of life; and we can clearly understand on this view the meaning of rudimentary organs. But disuse and selection will generally act on each creature, when it has