Origin of Species Variorum

Comparison with 1861

his work. But when many men, without intending to alter the breed, have a nearly common standard of perfection, and all try to get and breed from the best animals, much improvement and modification surely but slowly follow from this unconscious process of selection, notwithstanding a large amount of crossing with inferior animals. Thus it will be in nature; for within a confined area, with some place in its polity not so perfectly occupied as might be, natural selection will always tend to preserve all the individuals varying in the right direction, though in different degrees, so as better to fill up the unoccupied place. But if the area be large, its several districts will almost certainly present different conditions of life; and then if natural selection be modifying and improving a species in the several districts, there will be intercrossing with the other individuals of the same species on the confines of each. ↑ And in this case the effects of intercrossing can hardly be coun- terbalanced by natural selection always tending to modify all the individuals in each district in exactly the same manner to the conditions of each; for in a continuous area, the conditions will generally graduate away insensibly from one district to another. The intercrossing will most affect those animals which unite for each birth, which wander much, and which do not breed at a very quick rate. Hence in animals of this nature, for instance in birds, varieties will generally be confined to separated countries; and this I believe to be the case. In hermaphrodite organisms which cross only occasionally, and likewise in animals which unite for each birth, but which wander little and which can increase at a very rapid rate, a new and improved variety might be quickly formed on any one spot, and might there maintain itself in a body, so that whatever intercrossing took place would be chiefly between the individuals of the same new variety. A local variety when once thus formed might subsequently slowly spread to other districts. On the above principle, nurserymen

his work. But when many men, without intending to alter the breed, have a nearly common standard of perfection, and all try to procure and breed from the best animals, much improvement .. surely but slowly follows from this unconscious process of selection, notwithstanding a large amount of crossing with inferior animals. Thus it will be in nature; for within a confined area, with some place in its polity not .. perfectly occupied, natural selection will always tend to preserve all the individuals varying in the right direction, though in different degrees, so as better to fill up the unoccupied place. But if the area be very large, its several districts will almost certainly present different conditions of life; and then, ↑ if the same species undergoes modification in different parts, the newly-formed varieties will intercross on the confines of each district. But we shall see in the seventh chapter that intermediate varieties, inhabiting an intermediate district, whether the result of the crossing of other varieties, or originally formed with an intermediate character, will in the long run generally be supplanted by one of the varieties on either hand. .. Intercrossing will affect those animals most which unite for each birth and wander much, and which do not breed at a very quick rate. Hence with animals of this nature, for instance .. birds, varieties will generally be confined to separated countries; and this I find to be the case. With hermaphrodite organisms which cross only occasionally, and likewise with animals which unite for each birth, but which wander little and .. can increase at a very rapid rate, a new and improved variety might be quickly formed on any one spot, and might there maintain itself in a body and afterwards spread, so that the crossing would be chiefly between the individuals of the new variety living together in the same place. On this principle, nurserymen