Origin of Species Variorum

→ (E and F) which 1869

which 1859 1860 1861 1866

(E) and (F) which 1872

→ a 14 , 1859 1861 1866 1869 1872

a 14 1860

→ q 14 , 1859 1861 1866 1869 1872

, 1860

→ p 14 , 1859 1861 1866 1869 1872

q 14 , 1860

→ will 1859 1861 1866 1869 1872

p 14 , will 1860

→ a 10 ; 1859 1861 1866 1869 1872

a 10 1860

→ b 14 , 1861 1866 1869 1872

b 14 1859

; b 14 1860

→ f 14 , 1859 1861 1866 1869 1872

f 14 1860

→ a 5 , 1859 1861 1866 1869 1872

a 5 , 1860

→ o 14 , 1859 1861 1866 1869 1872

o 14 1860

→ e 14 , 1859 1861 1866 1869 1872

, e 14 , 1860

→ m 14 , 1859 1861 1866 1869 1872

m 14 1860

→ descendants. Hence the six new species descended from (I), and the eight descended from (A), will have to be ranked as very distinct genera, or even as distinct sub-families. 1869

descendants. 1859 1860 1861 1866 1872

original species will have transmitted offspring to the fourteen-thousandth generation. We may suppose that only one (F), of the two species →(E and F) which were least closely related to the other nine original species, has transmitted descendants to this late stage of descent.

The new species in our diagram descended from the original eleven species, will now be fifteen in number. Owing to the divergent tendency of natural selection, the extreme amount of difference in character between species a 14 and z 14 will be much greater than that between the most

different

distinct

of the original eleven species. The new species, moreover, will be allied to each other in a widely different

mauner.

manner.

Of the eight descendants from (A) the three marked → a 14 , → q 14 , → p 14 , →will be nearly related from having recently branched off from → a 10 ; → b 14 , and → f 14 ,

, from

from

having diverged at an earlier period from → a 5 , will be in some degree distinct from the three first-named species; and lastly, → o 14 , → e 14 , and → m 14 ,

, will

will

be nearly related one to the other,

but

but,

from having diverged at the first commencement of the process of modification, will be widely different from the other five species, and may constitute a sub-genus or

even

....

a distinct genus.

The six descendants from (I) will form two

subgenera

sub-genera

even

....

genera. But as the original species (I) differed largely from (A), standing nearly at the extreme

points

end

of the original genus, the six descendants from (I) will, owing to

inheritance,

inheritance

differ

alone, differ

considerably from the eight descendants from (A); the two groups, moreover, are supposed to have gone on diverging in different directions. The intermediate species, also (and this is a very important consideration), which connected the original species (A) and (I), have all become, excepting (F), extinct, and have left no →descendants. Hence the six new species descended from (I), and the eight descended from (A), will have to be ranked as very distinct genera, or even as distinct sub-families.