Origin of Species Variorum

quite new station, in which offspring and progenitor do not come into competition, both may continue to exist.

If, then, our diagram be assumed to represent a considerable amount of modification, species (A) and all the earlier varieties will have become extinct, being replaced by eight new species ( a 14 to m 14 ); and species (I) will be replaced by six ( n 14 to z 14 ) new species.

But we may go further than this. The original species of our genus were supposed to resemble each other in unequal degrees, as is so generally the case in nature; species (A) being more nearly related to B, C, and D, than to the other species; and species (I) more to G, H, K, L, than to the others. These two species (A) and (I) were also supposed to be very common and widely diffused species, so that they must originally have had some advantage over most of the other species of the genus. Their modified descendants, fourteen in number at the fourteen-thousandth generation, will probably have inherited some of the same advantages; they have also been modified and improved in a diversified manner at each stage of descent, so as to have become adapted to many related places in the natural economy of their country. It seems, therefore, .. extremely probable that they will have taken the places of, and thus exterminated, not only their parents (A) and (I), but likewise some of the original species which were most nearly related to their parents. Hence very few of the original species will have transmitted offspring to the fourteen-thousandth generation. We may suppose that only one (F), of the two species (E) and (F) which were least closely related to the other nine original species, has transmitted descendants to this late stage of descent.

The new species in our diagram descended from the original eleven species, will now be fifteen in number. Owing to the divergent tendency of natural selection, the extreme amount of difference in character between species a 14 and z 14 will be much greater than that between the most distinct of the original eleven species. The new species, moreover, will be allied to each other in a widely different manner. Of the eight descendants from (A) the three marked a 14 , q 14 , p 14 , will be nearly related from having recently branched off from a 10 ; b 14 , and f 14 , from having diverged at an earlier period from a 5 , will be in some degree distinct from the three first-named species; and lastly, o 14 , e 14 , and m 14 , will be nearly related one to the other, but, from having diverged at the first commencement of the process of modification, will be widely different from the other five species, and may constitute a sub-genus or .. a distinct genus.

The six descendants from (I) will form two sub-genera or .. genera. But as the original species (I) differed largely from (A), standing