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1859
1860
1861
1866
1869
1872

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1859
1860
1861
1866
1869

m 14 ); 1859 1861 1866 1869 1872
m 14 ); 1860

species (I) will be 1872
(I) will have been 1859 1860 1861 1866 1869

z 14 ) 1859 1861 1866 1869 1872
z 14 1860

(E) and (F) which 1872
which 1859 1860 1861 1866
(E and F) which 1869

a 14 , 1859 1861 1866 1869 1872
a 14 1860

q 14 , 1859 1861 1866 1869 1872
, 1860

p 14 , 1859 1861 1866 1869 1872
q 14 , 1860

will 1859 1861 1866 1869 1872
p 14 , will 1860

a 10 ; 1859 1861 1866 1869 1872
a 10 1860

b 14 , 1861 1866 1869 1872
b 14 1859
; b 14 1860

f 14 , 1859 1861 1866 1869 1872
f 14 1860

a 5 , 1859 1861 1866 1869 1872
a 5 , 1860

o 14 , 1859 1861 1866 1869 1872
o 14 1860

e 14 , 1859 1861 1866 1869 1872
, e 14 , 1860

m 14 , 1859 1861 1866 1869 1872
m 14 1860

quite new station, in which
child
offspring
and
parent
progenitor
do not come into competition, both may continue to exist.
If
If,
then
then,
our diagram be assumed to represent a considerable amount of modification, species (A) and all the earlier varieties will have become extinct,
having been
being
replaced by eight new species ( a 14 to m 14 ); and species (I) will be replaced by six ( n 14 to z 14 )
) new
new
species.
But we may go further than this. The original species of our genus were supposed to resemble each other in unequal degrees, as is so generally the case in nature; species (A) being more nearly related to B, C, and D, than to the other species; and species (I) more to G, H, K, L, than to the others. These two species (A) and
(I),
(I)
were also supposed to be very common and widely diffused species, so that they must originally have had some advantage over most of the other species of the genus. Their modified descendants, fourteen in number at the fourteen-thousandth generation, will probably have inherited some of the same
advantages:
advantages;
they have also been modified and improved in a diversified manner at each stage of descent, so as to have become adapted to many related places in the natural economy of their country. It seems, therefore,
to me
....
extremely probable that they will have taken the places of, and thus exterminated, not only their parents (A) and (I), but likewise some of the original species which were most nearly related to their parents. Hence very few of the original species will have transmitted offspring to the fourteen-thousandth generation. We may suppose that only one (F), of the two species (E) and (F) which were least closely related to the other nine original species, has transmitted descendants to this late stage of descent.
The new species in our diagram descended from the original eleven species, will now be fifteen in number. Owing to the divergent tendency of natural selection, the extreme amount of difference in character between species a 14 and z 14 will be much greater than that between the most
different
distinct
of the original eleven species. The new species, moreover, will be allied to each other in a widely different
mauner.
manner.
Of the eight descendants from (A) the three marked a 14 , q 14 , p 14 , will be nearly related from having recently branched off from a 10 ; b 14 , and f 14 ,
, from
from
having diverged at an earlier period from a 5 , will be in some degree distinct from the three first-named species; and lastly, o 14 , e 14 , and m 14 ,
, will
will
be nearly related one to the other,
but
but,
from having diverged at the first commencement of the process of modification, will be widely different from the other five species, and may constitute a sub-genus or
even
....
a distinct genus.
The six descendants from (I) will form two
subgenera
sub-genera
or
even
....
genera. But as the original species (I) differed largely from (A), standing