Origin of Species Variorum

Comparison with 1861

all such valuations being at present empirical. ↑ Moreover, on the view of the origin of genera which I shall presently give, we have no right to expect often to meet with generic differences in our domesticated productions.

When we attempt to estimate the amount of structural difference between the domestic races of the same species, we are soon involved in doubt, from not knowing whether they have descended from one or several parent-species. This point, if it could be cleared up, would be interesting; if, for instance, it could be shown that the greyhound, bloodhound, terrier, spaniel, and bull-dog, which we all know propagate their kind so truly, were the offspring of any single species, then such facts would have great weight in making us doubt about the immutability of the many very closely allied and natural species — for instance, of the many foxes — inhabiting different quarters of the world. I do not believe, as we shall presently see, that ↑ the whole amount of difference between the several breeds of the dog has been produced under domestication; I believe that some small part of the difference is due to their having descended from distinct species. In the case of some other domesticated species, there is presumptive, or even strong evidence, that all the breeds have descended from a single wild stock.

It has often been assumed that man has chosen for domestication animals and plants having an extraordinary inherent tendency to vary, and likewise to withstand diverse climates. I do not dispute that these capacities have added largely to the value of most of our domesticated productions; but how could a savage possibly know, when he first tamed an animal, whether it would vary in succeeding generations, and whether it would endure other climates? Has the little variability of the ass or guinea-fowl, or the small power of endurance

all such valua- tions being at present empirical. When it is explained how genera originate under nature, it will be seen that we have no right to expect often to find a generic amount of difference in our domesticated races. ↑

In attempting to estimate the amount of structural difference between the domestic races of the same species, we are soon involved in doubt, from not knowing whether they have descended from one or several parent-species. This point, if it could be cleared up, would be interesting; if, for instance, it could be shown that the greyhound, bloodhound, terrier, spaniel, and bull-dog, which we all know propagate their kind so truly, were the offspring of any single species, then such facts would have great weight in making us doubt about the immutability of the many very closely allied .. natural species — for instance, of the many foxes — inhabiting different quarters of the world. I do not believe, as we shall presently see, that the whole amount of difference between the several breeds of the dog has been produced under domestication; I believe that some small part of the difference is due to their having descended from distinct species. In the case of strongly marked races in some other domesticated species, there is presumptive, or even strong evidence, that all are descended from a single wild stock.