Origin of Species Variorum

Comparison with 1859

it, nature is prodigal in variety, but niggard in innovation. Why, on the theory of Creation, should this be so? Why should all the parts and organs of many independent beings, each supposed to have been separately created for its proper place in nature, be so invariably linked together by graduated steps? Why should not Nature have taken a leap from structure to structure? On the theory of natural selection, we can clearly understand why she should not; for natural selection can act only by taking advantage of slight successive variations; she can never take a leap, but must advance by the shortest and slowest steps.

Organs of little apparent importance .—

As natural selection acts by life and death,— by the preservation of individuals with any favourable variation, and by the destruction of those with any unfavourable deviation of structure,— I have sometimes felt much difficulty in understanding the origin of simple parts, of which the importance does not seem sufficient to cause the preservation of successively varying individuals. I have sometimes felt as much difficulty, though of a very different kind, on this head, as in the case of an organ as perfect and complex as the eye.

In the first place, we are much too ignorant in regard to the whole economy of any one organic being, to say what slight modifications would be of importance or not. In a former chapter I have given instances of most trifling characters, such as the down on fruit and the colour of the flesh, which, from determining the attacks of insects or from being correlated with constitutional differences, might assuredly be acted on by natural selection. The tail of the giraffe looks like an artificially constructed fly-flapper; and it seems at first incredible that this could have been adapted for its present purpose

it, Nature is prodigal in variety, but niggard in innovation. Why, on the theory of Creation, should there be so much variety and so little novelty? Why should all the parts and organs of many independent beings, each supposed to have been separately created for its proper place in nature, be so commonly linked together by graduated steps? Why should not Nature .. take a sudden leap from structure to structure? On the theory of natural selection, we can clearly understand why she should not; for natural selection .. acts only by taking advantage of slight successive variations; she can never take a sudden leap, but must advance by short and sure though slow steps.

Organs of little apparent Importance, as affected by Natural Selection.

As natural selection acts by life and death,—by .. the survival of the fittest, and by the destruction of the less well fitted individuals,—I have sometimes felt great difficulty in understanding the origin or formation of parts of little importance; almost as great, though of a very different kind, as in the case of the most perfect and complex organs. ↑

In the first place, we are much too ignorant in regard to the whole economy of any one organic being, to say what slight modifications would be of importance or not. In a former chapter I have given instances of very trifling characters, such as the down on fruit and the colour of its flesh, the colour of the skin and hair of quadrupeds, which, from being correlated with constitutional differences or from determining the attacks of insects, might assuredly be acted on by natural selection. The tail of the giraffe looks like an artificially constructed fly-flapper; and it seems at first incredible that this could have been adapted for its present pur- pose