Origin of Species Variorum

Comparison with 1866

convinced that a damp climate affects the growth of the hair, and that with the hair the horns are correlated. Mountain breeds always differ from lowland breeds; and a mountainous country would probably affect the hind limbs from exercising them more, and possibly even the form of the pelvis; and then by the law of homologous variation, the front limbs and even the head would probably be affected. The shape, also, of the pelvis might affect by pressure the shape of the head of the young in the womb. The laborious breathing necessary in high regions would, we have some reason to believe, increase the size of the chest; and again correlation would come into play. The effects on the whole organisation of lessened exercise with abundant food is probably still more important; and this, as H. von Nathusius has lately shown in his excellent Treatise, is apparently one chief cause of the great modification which the breeds of swine have undergone. Animals kept by savages in different countries often have to struggle for their own subsistence, and would be exposed to a certain extent to natural selection, and individuals with slightly different constitutions would succeed best under different climates; and there is reason to believe that constitution and colour are correlated. A good observer, also, states that in cattle susceptibility to the attacks of flies is correlated with colour, as is the liability to be poisoned by certain plants; so that colour would be thus subjected to the action of natural selection. But we are far too ignorant to speculate on the relative importance of the several known and unknown laws of variation; and I have here alluded to them only to show that, if we are unable to account for the characteristic differences of our domestic breeds, which nevertheless we generally admit to have arisen through ordinary generation, we ought not to lay too much stress on our ignorance of the precise cause of the slight analogous differences between species. I might have adduced for this same purpose the differences between the races of man, which are so strongly marked; I may add that some little light can apparently be thrown on the origin of these differences, chiefly through sexual selection of a particular kind, but without here entering on copious

convinced that a damp climate affects the growth of the hair, and that with the hair the horns are correlated. Mountain breeds always differ from lowland breeds; and a mountainous country would probably affect the hind limbs from exercising them more, and possibly even the form of the pelvis; and then by the law of homologous variation, the front limbs and .. the head would probably be affected. The shape, also, of the pelvis might affect by pressure the shape of certain parts of the young in the womb. The laborious breathing necessary in high regions would, we have .. reason to believe, increase the size of the chest; and again correlation would come into play. The effects on the whole organisation of lessened exercise together with abundant food is probably still more important; and this, as H. von Nathusius has lately shown in his excellent Treatise, is apparently one chief cause of the great modification which the breeds of swine have undergone. Animals kept by savages in different countries often have to struggle for their own subsistence, and are exposed to a certain extent to natural selection, and individuals with slightly different constitutions would succeed best under different climates. .. .. .. .. .. .. .. .. .. .. .. .. A good observer .. states that in cattle susceptibility to the attacks of flies is correlated with colour, as is the liability to be poisoned by certain plants; so that even colour would be thus subjected to the action of natural selection. But we are far too ignorant to speculate on the relative importance of the several known and unknown causes of variation; and I have made these remarks only to show that, if we are unable to account for the characteristic differences of our domestic breeds, which nevertheless are generally admitted to have arisen through ordinary generation from one or a few parent-stocks, we ought not to lay too much stress on our ignorance of the precise cause of the slight analogous differences between species. I might have adduced for this same purpose the differences between the races of man, which are so strongly marked; I may add that some .. light can apparently be thrown on ... these differences, .. through sexual selection of a particular kind, but without .. entering on full