Origin of Species Variorum

Comparison with 1859

the productions on the shores of a continent when first broken up into an archipelago), and consequently during subsidence, though there will be much extinction, fewer new varieties or species will be formed; and it is during these very periods of subsidence, that our great deposits rich in fossils have been accumulated. Nature may almost be said to have guarded against the frequent discovery of her transitional or linking forms. ↑

From the foregoing considerations it cannot be doubted that the geological record, viewed as a whole, is extremely imperfect; but if we confine our attention to any one formation, it becomes more difficult to under- stand, why we do not therein find closely graduated varieties between the allied species which lived at its commencement and at its close. Some cases are on record of the same species presenting distinct varieties in the upper and lower parts of the same formation, but, as they are rare, they may be here passed over. Although each formation has indisputably required a vast number of years for its deposition, I can see several reasons why each should not include a graduated series of links between the species which then lived; but I can by no means pretend to assign due proportional weight to the following considerations.

Although each formation may mark a very long lapse of years, each perhaps is short compared with the period requisite to change one species into another. I am aware that two palæontologists, whose opinions are worthy of much deference, namely Bronn and Woodward,

.. on the shores of a continent when first broken up into an archipelago), and consequently during subsidence, though there will be much extinction, few new varieties or species will be formed; and it is during these very periods of subsidence, that the deposits which are richest in fossils have been accumulated. ↑

On the Absence of Numerous Intermediate Varieties in any one Single Formation .

From these several considerations, it cannot be doubted that the geological record, viewed as a whole, is extremely imperfect; but if we confine our attention to any one formation, it becomes much more difficult to understand why we do not therein find closely graduated varieties between the allied species which lived at its commencement and at its close. Several cases are on record of the same species presenting .. varieties in the upper and lower parts of the same formation: thus, Trautschold gives a number of instances with Ammonites; and Hilgendorf has described a most curious case of ten graduated forms of Planorbis multiformis in the successive beds of a fresh-water formation in Switzerland. Although each formation has indisputably required a vast number of years for its deposition, ... several reasons can be given why each should not commonly include a graduated series of links between the species which .. lived at its commencement and close; but I cannot assign due proportional weight to the following considerations.

Although each formation may mark a very long lapse of years, each probably is short compared with the period requisite to change one species into another. I am aware that two palæontologists, whose opinions are worthy of much deference, namely Bronn and Wood- ward,