Origin of Species Variorum

Comparison with 1859

on the bed of the sea, or where it did not accumulate at a sufficient rate to protect organic bodies from decay, no remains could be preserved.

In our archipelago, I believe that fossiliferous formations could be formed of sufficient thickness to last to an age, as distant in futurity as the secondary formations lie in the past, only during periods of subsidence. These periods of subsidence would be separated from each other by enormous intervals, during which the area would be either stationary or rising; whilst rising, each fossiliferous formation would be destroyed, almost as soon as accumulated, by the incessant coast-action, as we now see on the shores of South America. ↑ During the periods of subsidence there would probably be much extinction of life; during the periods of elevation, there would be much variation, but the geological record would then be least perfect.

It may be doubted whether the duration of any one great period of subsidence over the whole or part of the archipelago, together with a contemporaneous accumulation of sediment, would exceed the average duration of the same specific forms; and these contingencies are indispensable for the preservation of all the transitional gradations between any two or more species. If such gradations were not fully preserved, transitional varieties would merely appear as so many distinct species. It is, also, probable that each great period of subsidence would be interrupted by oscillations

on the bed of the sea, or where it did not accumulate at a sufficient rate to protect organic bodies from decay, no remains could be preserved.

Formations rich in fossils of many kinds, and of thickness sufficient to last to an age as distant in futurity as the secondary formations lie in the past, would generally be formed in the archipelago only during periods of subsidence. These periods of subsidence would be separated from each other by immense intervals of time, during which the area would be either stationary or rising; whilst rising, the fossiliferous formations on the steeper shores would be destroyed, almost as soon as accumulated, by the incessant coast-action, as we now see on the shores of South America; even throughout the extensive and shallow seas within the archipelago sedimentary beds could hardly be accumulated of great thickness during the periods of elevation, or become capped and protected by subsequent deposits, so as to have a good chance of enduring to a very distant future. ↑ During the periods of subsidence, there would probably be much extinction of life; during the periods of elevation, there would be much variation, but the geological record would then be least perfect.

It may be doubted whether the duration of any one great period of subsidence over the whole or part of the archipelago, together with a contemporaneous accumulation of sediment, would exceed the average duration of the same specific forms; and these contingencies are indispensable for the preservation of all the transitional gradations between any two or more species. If such gradations were not all fully preserved, transitional varieties would merely appear as so many new and distinct species. It is also probable that each great period of subsidence would be interrupted by oscillations