the more important it becomes for classification. As an instance: Owen, in speaking of the dugong, says, "The generative
organs, organs, 1866 1869 1872 | organs 1859 1860 1861 |
being those which are most remotely related to the habits and food of an animal, I have always regarded as affording very clear indications of its true affinities. We are least likely in the modifications of these organs to mistake a merely adaptive for an essential character."
....... 1869 1872 | So 1859 1860 1861 1866 |
With With 1869 1872 | with 1859 1860 1861 1866 |
plants, plants, 1859 1860 1861 1866 1869 | plants 1872 |
how remarkable it is that the organs of vegetation, on which their
nutrition and nutrition and 1869 1872 | whole 1859 1860 1861 1866 |
life
depend, depend, 1869 1872 | depends, 1859 1860 1861 1866 |
are of little
signification; signification; 1869 1872 | signification, 1859 1860 1861 1866 |
...OMIT 1869 1872 |
excepting in the first main divisions; 1859 1860 1861 1866 |
whereas the organs of reproduction, with their product the
seed seed 1869 1872 | seed, 1859 1860 1861 1866 |
and embryo, are and embryo, are 1869 1872 |
are 1859 1860 1861 1866 |
of paramount importance! So again in formerly discussing
..
morphological
differences differences 1869 | characters 1872 |
which are not
physiologically physiologically 1869 | functionally 1872 |
important, we have seen that they are often of the highest service in classification. This depends on their constancy throughout many allied groups; and
the
constancy
depends chiefly depends chiefly 1869 | chiefly depends 1872 |
on any slight deviations
of structure in such parts not of structure in such parts not 1869 |
not 1872 |
having been preserved and accumulated by natural selection, which acts only on
useful useful 1869 | serviceable 1872 |
characters. ↑4 blocks not present in 1869 1872; present in 1859 1860 1861 1866 | We must not, therefore, in classifying, trust to resemblances in parts of the organisation, however important they may be for the welfare of the being in relation to the outer world.
Perhaps from this cause it has partly arisen, that almost all naturalists lay the greatest stress on resemblances in organs of high vital or physiological importance.
No doubt this view of the classificatory importance of organs which are important is generally, but by no means always, true.
But their importance for classification, I believe, depends on their greater constancy throughout large groups of species; and this constancy depends on such organs having generally been subjected to less change in the adaptation of the
species to their conditions of life.
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↑7 blocks not present in 1869; present in 1859 1860 1861 1866 1872 | That the mere physiological importance of an organ does not determine its classificatory value, is almost shown
by the one
fact, that in allied groups, in which the same organ, as we have every reason to suppose, has nearly the same physiological value, its classificatory value is widely different.
No naturalist can have worked at
any group without being struck with this fact; and it has been most
fully acknowledged in the writings of almost every author.
It will suffice to quote the highest authority, Robert Brown, who
in speaking of certain organs in the Proteaceæ,
says their generic importance, "like that of all their parts, not only in this
but, as I apprehend, in every natural family, is very unequal, and in some cases seems to be entirely lost."
Again
in another work he says, the genera of the Connaraceæ "differ in having one or more ovaria, in the existence or absence of albumen, in the imbricate or valvular æstivation.
Any one of these characters singly is frequently of more than generic importance, though here even when all taken together they appear insufficient to separate Cnestis from Connarus."
To give an example amongst insects,
in one great division of the Hymenoptera, the antennæ, as Westwood has remarked, are most constant in structure; in another division they differ much,
and the differences are of quite subordinate value in classification; yet no one probably
will say that the antennæ in these two divisions of the same order are of unequal physiological importance.
Any number of instances could be given of the varying importance for classification of the same important organ within the same group of beings.
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That the mere physiological importance of an organ does not determine its classificatory value, is almost proved by the fact, that in allied groups, in which the same organ, as we have every reason to suppose, has nearly the same physiological value, its classificatory value is widely different. No naturalist can have worked at any group without being struck with this fact; and it has been fully acknowledged in the writings of almost every author. It will suffice to quote the highest authority, Robert Brown, who, in speaking of certain organs in the Proteaceæ, says their generic importance, "like that of all their parts, not only in this, but, as I apprehend, in every natural family, is very unequal, and in some cases seems to be entirely lost." Again, in another work he says, the genera of the Connaraceæ
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