Origin of Species Variorum

Comparison with 1861

Would it be thought sufficient to say that because planets revolve in elliptic courses round the sun, satellites follow the same course round the planets, for the sake of symmetry, and to complete the scheme of nature? An eminent physiologist accounts for the presence of rudimentary organs, by supposing that they serve to excrete matter in excess, or injurious to the system; but can we suppose that the minute papilla; which often represents the pistil in male flowers, and which is formed merely of cellular tissue, can thus act? Can we suppose that the formation of rudimentary teeth which are subsequently absorbed, can be of any service to the rapidly growing embryonic calf by the excretion of precious phosphate of lime? When a man's fingers have been amputated, imperfect nails sometimes appear on the stumps: I could as soon believe that these vestiges of nails have appeared, not from unknown laws of growth, but in order to excrete horny matter, as that the rudimentary nails on the fin of the manatee were formed for this purpose.

On my view of descent with modification, the origin of rudimentary organs is simple. We have plenty of cases of rudimentary organs in our domestic productions,— as the stump of a tail in tailless breeds,— the vestige of an ear in earless breeds,— the reappearance of minute dangling horns in hornless breeds of cattle, more especially, according to Youatt, in young animals,— and the state of the whole flower in the cauliflower. We often see rudiments of various parts in monsters. But I doubt whether any of these cases throw light on the origin of rudimentary organs in a state of nature, further than by showing that rudiments can be produced; for I doubt whether species under nature ever undergo abrupt changes. I believe that disuse has been the main agency; that it has led in successive generations to the gradual reduction of various

bones? What would be thought of an astronomer, who maintained that the satellites revolve in elliptic courses round their planets "for the sake of symmetry," because the planets thus revolve round the sun? An eminent physiologist accounts for the presence of rudimentary organs, by supposing that they serve to excrete matter in excess, or matter injurious to the system; but can we suppose that the minute papilla, which often represent the pistil in male flowers, and which is formed .. of mere cellular tissue, can thus act? Can we suppose that ... rudimentary teeth, which are subsequently absorbed, are beneficial to the rapidly growing embryonic calf by removing matter so precious as phosphate of lime? When a mans fingers have been amputated, imperfect nails have been known to appear on the stumps, and I could as soon believe that these vestiges of nails have been developed in order to excrete horny matter, as that the rudimentary nails on the fin of the manatee have been developed for this same purpose.

On the view of descent with modification, the origin of rudimentary organs is simple. We have plenty of cases of rudimentary organs in our domestic productions,— as the stump of a tail in tailless breeds,— the vestige of an ear in earless breeds of sheep,— the reappearance of minute dangling horns in hornless breeds of cattle, more especially, according to Youatt, in young animals,— and the state of the whole flower in the cauliflower. We often see rudiments of various parts in monsters. But I doubt whether any of these cases throw light on the origin of rudimentary organs in a state of nature, further than by showing that rudiments can be produced; for I doubt whether species under nature ever undergo abrupt changes. I believe that disuse has been the main agency; that it has led in successive generations to the gradual reduction of various