Origin of Species Variorum

Comparison with 1866

under variable and under local forms, under geographical races or sub-species, and as true representative species. The variable forms vary much within the limits of the same island. The local forms of the same species are moderately constant and distinct in each separate island; but when all such forms from the several islands are compared, the differences become so slight, numerous, and graduated, that it is impossible to define or describe many of them, though at the same time the extreme forms are sufficiently distinct. The geographical races or sub-species are local forms completely fixed and isolated; but as they do not differ from each other by strongly marked and important characters, "there is no possible test but individual opinion to determine which of them shall be considered as species and which as varieties." Lastly, representative species fill the same place in the natural economy of each island as do the local forms and sub-species; but as they are distinguished from each other by a greater, though not by a definite, amount of difference than that between the local forms and sub-species, they are almost universally ranked by naturalists as true species. Nevertheless, no certain criterion can possibly be given by which variable forms, local forms, sub-species, and representative species can be recognised. Many years ago, when comparing, and seeing others compare, the birds from the separate islands of the Galapagos Archipelago, both one with another, and with those from the American mainland, I was much struck how entirely vague and arbitrary is the distinction between species and varieties. On the islets of the little Madeira group there are many insects which are characterized as varieties in Mr. Wollaston's admirable work, but which certainly would be ranked as distinct species by many entomologists. Even Ireland

under four heads, namely, as variable forms, as local forms, as geographical races or sub-species, and as true representative species. The first or variable forms vary much within the limits of the same island. The local forms ... are moderately constant and distinct in each separate island; but when all the forms from the several islands are compared, the differences are seen to be so slight .. and graduated, that it is impossible to define or describe .. them, though at the same time the extreme forms are sufficiently distinct. The geographical races or sub-species are local forms completely fixed and isolated; but as they do not differ from each other by strongly marked and important characters, "there is no possible test but individual opinion to determine which of them shall be considered as species and which as varieties." Lastly, representative species fill the same place in the natural economy of each island as do the local forms and sub-species; but as they are distinguished from each other by a greater ... amount of difference than that between the local forms and sub-species, they are almost universally ranked by naturalists as true species. Nevertheless, no certain criterion can possibly be given by which variable forms, local forms, sub-species, and representative species can be recognised. Many years ago, when comparing, and seeing others compare, the birds from the closely neighbouring islands of the Galapagos archipelago, both one with another, and with those from the American mainland, I was much struck how entirely vague and arbitrary is the distinction between species and varieties. On the islets of the little Madeira group there are many insects which are characterized as varieties in Mr. Wollaston's admirable work, but which would certainly be ranked as distinct species by many entomologists. Even Ire- land