Origin of Species Variorum

alliance of many others, on the most distant mountains, and in the northern and southern temperate zones; and likewise the close alliance of some of the inhabitants of the sea in the northern and southern temperate latitudes, though separated by the whole intertropical ocean. Although two countries may present .. physical conditions as closely similar as the same species ever require, we need feel no surprise at their inhabitants being widely different, if they have been for a long period completely sundered from each other; for as the relation of organism to organism is the most important of all relations, and as the two countries will have received colonists ... at various periods and in different proportions, from some other country or from each other, the course of modification in the two areas will inevitably have been different.

On this view of migration, with subsequent modification, we .. see why oceanic islands are inhabited by only few species, but of these, why many are peculiar or endemic forms. We .. clearly see why species belonging to those groups of animals which cannot cross wide spaces of the ocean, as frogs and terrestrial mammals, do not inhabit oceanic islands; and why, on the other hand, new and peculiar species of bats, animals which can traverse the ocean, are so often found on islands far distant from any continent. Such cases as the presence of peculiar species of bats on oceanic islands, and the absence of all other terrestrial mammals, are facts utterly inexplicable on the theory of independent acts of creation.

The existence of closely allied or representative species in any two areas, implies, on the theory of descent with modification, that the same parent-forms formerly inhabited both areas; and we almost invariably find that wherever many closely allied species inhabit two