Origin of Species Variorum

Comparison with 1866

of its range, where it exists in lessened numbers, will, during fluctuations in the number of its enemies or of its prey, or in the seasons, be extremely liable to utter extermination; and thus its geographical range will come to be still more sharply defined.

If I am right in believing that allied or representative species, when inhabiting a continuous area, are generally so distributed that each has a wide range, with a comparatively narrow neutral territory between them, in which they become rather suddenly rarer and rarer; then, as varieties do not essentially differ from species, the same rule will probably apply to both; and if we in imagination adapt a varying species to a very large area, we shall have to adapt two varieties to two large areas, and a third variety to a narrow intermediate zone. The intermediate variety, consequently, will exist in lesser numbers from inhabiting a narrow and lesser area; and practically, as far as I can make out, this rule holds good with varieties in a state of nature. I have met with striking instances of the rule in the case of varieties intermediate between well-marked varieties in the genus Balanus. And it would appear from information given me by Mr. Watson, Dr. Asa Gray, and Mr. Wollaston, that generally when varieties intermediate between two other forms occur, they are much rarer numerically than the forms which they cannect. Now, if we may trust these facts and inferences, and therefore conclude that varieties linking two other varieties together have generally existed in lesser numbers than the forms which they connect, then, I think, we can understand why intermediate varieties should not endure for very long periods;—why, .. as a general rule they should be exterminated and disappear, sooner than the forms which they originally linked together.

For any form existing in lesser numbers would, as already remarked, run a greater chance of being exterminated than one existing in large numbers; and in this particular case the intermediate form would be eminently liable to the inroads of closely allied forms existing on both sides of it. But a far more important consideration, as I believe, is that, during the process of further modification, by which two varieties are supposed on my theory to be converted and perfected into two distinct species, the two which exist in larger numbers from inhabiting larger areas, will have a great advantage over the intermediate variety, which exists in smaller numbers in a narrow and intermediate zone. For forms existing in larger numbers will always have a better chance, within any given period, of presenting further favourable variations for natural selection to seize on, than will the rarer forms which exist in lesser numbers. Hence, the more common forms, in the race for life, will tend to beat and supplant the less common forms, for these will be more

of its range, where it exists in lessened numbers, will, during fluctuations in the number of its enemies or of its prey, or in the nature of the seasons, be extremely liable to utter extermination; and thus its geographical range will come to be still more sharply defined.

As allied or representative species, when inhabiting a continuous area, are generally .. distributed in such a manner that each has a wide range, with a comparatively narrow neutral territory between them, in which they become rather suddenly rarer and rarer; then, as varieties do not essentially differ from species, the same rule will probably apply to both; and if we take a varying species inhabiting a very large area, we shall have to adapt two varieties to two large areas, and a third variety to a narrow intermediate zone. The intermediate variety, consequently, will exist in lesser numbers from inhabiting a narrow and lesser area; and practically, as far as I can make out, this rule holds good with varieties in a state of nature. I have met with striking instances of the rule in the case of varieties intermediate between well-marked varieties in the genus Balanus. And it would appear from information given me by Mr. Watson, Dr. Asa Gray, and Mr. Wollaston, that generally, when varieties intermediate between two other forms occur, they are much rarer numerically than the forms which they connect. Now, if we may trust these facts and inferences, and .. conclude that varieties linking two other varieties together generally have existed in lesser numbers than the forms which they connect, then .. we can understand why intermediate varieties should not endure for very long periods:— why, as a general rule, they should be exterminated and disappear, sooner than the forms which they originally linked together.

For any form existing in lesser numbers would, as already remarked, run a greater chance of being exterminated than one existing in large numbers; and in this particular case the intermediate form would be eminently liable to the inroads of closely-allied forms existing on both sides of it. But it is a far more important consideration, ... that during the process of further modification, by which two varieties are supposed ... to be converted and perfected into two distinct species, the two which exist in larger numbers, from inhabiting larger areas, will have a great advantage over the intermediate variety, which exists in smaller numbers in a narrow and intermediate zone. For forms existing in larger numbers will .. have a better chance, within any given period, of presenting further favourable variations for natural selection to seize on, than will the rarer forms which exist in lesser numbers. Hence, the more common forms, in the race for life, will tend to beat and supplant the less common forms, for these will be more