Origin of Species Variorum

Comparison with 1869

constructed is the feathered wing of a bird and the membrane-covered wing of a bat with all the digits largely developed; and still more so the four wings of a butterfly, the two wings of a fly, and the two wings of a beetle, together with the elytra. Bivalve shells have only to open and shut, but on what a number of patterns is the hinge constructed, from the long row of neatly interlocking teeth in a Nucula to the simple ligament of a Mussel. Seeds are disseminated by their minuteness,—by their capsule being converted into a light ballon-like envelope,—by being embedded in pulp or flesh, formed of the most diverse parts, and rendered nutritious as well as conspicuously coloured, so as to attract and be devoured by birds,—by having hooks and grapnels of many kinds and serrated awns, so as to adhere to the fur of quadrupeds,—and by being furnished with wings and plumes, as diversified in shape as elegant in structure, so as to be wafted by every breeze. I will give one other instance; for the subject is worthy of reflection by those who are not able to credit that organic beings have been formed in many ways for the sake of mere variety, like toys in a shop. Some authors maintain that organic beings have been formed in many ways for the sake of mere variety, almost like toys in a shop, but such a view of nature is incredible. With plants having separated sexes, and with those in which, though hermaphrodites, the pollen does not spontaneously fall on the stigma, some aid is necessary for their fertilisation. With several kinds this is effected by the light and incoherent pollen-grains being blown by the wind through mere chance on to the stigma; and this is the simplest plan which can well be conceived. An almost equally simple, though very different, plan occurs in many cases, in which a symmetrical flower secretes a few drops of nectar, and is consequently visited by insects; and these carry the pollen from the anthers to the stigma.

From this simple stage we may pass through an in-exhaustible number of contrivances, all for the same purpose and effected in essentially the same manner, but entailing changes in every part of the flower; with the nectar stored in variously shaped receptacles, with the stamens and pistils modified in many ways, sometimes forming trap-like contrivances, and sometimes capable of neatly adapted movements through irritability or elasticity. From such structures we may advance till we come to such an acme of perfect adaptation, as has lately been described by Dr. Crüger in the case of Coryanthes. This orchid has its labellum or lower lip hollowed out into a great bucket, into which drops of almost pure water, not nectar, continually fall from two secreting horns which stand above it; and when the bucket is half full, the water overflows by a spout on one side. The basal part of the labellum curves over the bucket, and is itself hollowed out into a sort of chamber with two

constructed is the feathered wing of a bird and the membrane-covered wing of a bat; ... and still more so the four wings of a butterfly, the two wings of a fly, and the two wings with the elytra of a beetle. Bivalve shells are made to open and shut, but on what a number of patterns is the hinge constructed,— from the long row of neatly interlocking teeth in a Nucula to the simple ligament of a Mussel! Seeds are disseminated by their minuteness,— by their capsule being converted into a light balloon-like envelope,— by being embedded in pulp or flesh, formed of the most diverse parts, and rendered nutritious, as well as conspicuously coloured, so as to attract and be devoured by birds,— by having hooks and grapnels of many kinds and serrated awns, so as to adhere to the fur of quadrupeds,— and by being furnished with wings and plumes, as different in shape as they are elegant in structure, so as to be wafted by every breeze. I will give one other instance; for this subject of the same end being gained by the most diversified means well deserves attention. Some authors maintain that organic beings have been formed in many ways for the sake of mere variety, almost like toys in a shop, but such a view of nature is incredible. With plants having separated sexes, and with those in which, though hermaphrodites, the pollen does not spontaneously fall on the stigma, some aid is necessary for their fertilisation. With several kinds this is effected by the pollen-grains, which are light and incoherent, .. being blown by the wind through mere chance on to the stigma; and this is the simplest plan which can well be conceived. An almost equally simple, though very different, plan occurs in many plants in which a symmetrical flower secretes a few drops of nectar, and is consequently visited by insects; and these carry the pollen from the anthers to the stigma.

From this simple stage we may pass through an inexhaustible number of contrivances, all for the same purpose and effected in essentially the same manner, but entailing changes in every part of the flower. The nectar may be stored in variously shaped receptacles, with the stamens and pistils modified in many ways, sometimes forming trap-like contrivances, and sometimes capable of neatly adapted movements through irritability or elasticity. From such structures we may advance till we come to such a case of extraordinary adaptation as that lately .. described by Dr. Crüger in the .. Coryanthes. This orchid has part of its labellum or lower lip hollowed out into a great bucket, into which drops of almost pure water .. continually fall from two secreting horns which stand above it; and when the bucket is half full, the water overflows by a spout on one side. The basal part of the labellum stands over the bucket, and is itself hollowed out into a sort of chamber with two