See page in:
1859
1860
1861
1866
1869
1872

Compare with:
1859
1860
1861
1866
1869

with its beak 1872
away 1861 1866 1869

all the slight individual variations in the shape of the beak, which were 1869 1872
each slight variation of beak, 1861 1866

or spontaneous 1869 1872
from the want of other food, or the preservation of chance 1861 1866

to become more 1869 1872
more 1861 1866

or taste; but 1869 1872
but 1861 1866

it was instinct 1869 1872
instinct 1861 1866

varied. 1869 1872
slightly changed; nor can we conjecture by what gradations many instincts have been developed when they relate to organs (such as the mammary glands) on the first origin of which we know nothing. 1861 1866

were independently acquired through 1869 1872
have been acquired by independent acts of 1859 1860 1861
have been independently acquired by 1866

this bird often holds the seeds of the yew between its feet on a branch, and hammers with its beak till it gets
into
at
the kernel. Now what special difficulty would there be in natural selection preserving all the slight individual variations in the shape of the beak, which were better and better adapted to break open
seeds,
the seeds,
until a beak was formed, as well constructed for this purpose as that of the nuthatch, at the same time that
hereditary
....
habit, or
compulsion
compulsion,
or spontaneous variations of taste,
made
led
the bird to become more and more of a seed-eater? In this case the beak is supposed to be slowly modified by natural selection, subsequently to, but in accordance with, slowly changing
habit;
habits
or taste; but let the feet of the titmouse vary and grow larger from correlation with the beak, or from any other unknown cause, and
is
....
it
very
is not
improbable that such larger feet
might
would
lead the bird to climb more
and more
and more
until it acquired
even
....
the remarkable climbing instinct and
capacity
power
of the
nuthatch?
nuthatch.
In this case a gradual change of structure is supposed to lead to changed instinctive
habits
habits.
of
....
life.
....
To take one more case: few instincts are more remarkable than that which leads the swift of the Eastern Islands to make its nest wholly of inspissated saliva. Some birds build their nests of mud, believed to be moistened with saliva; and one of the swifts of North America makes its nest (as I have seen) of sticks agglutinated with saliva, and even with flakes of this substance. Is it then very improbable that the natural selection of individual swifts, which secreted more and more saliva, should at last produce a species with instincts leading it to neglect other materials, and to make its nest exclusively of inspissated saliva? And so in other cases. It
must
must,
be
however, be
admitted that in many instances we cannot conjecture whether it was instinct or structure
has
which
first varied.
No doubt many instincts of very difficult explanation could be opposed to the theory of natural
selection,—
selection—
cases, in which we cannot see how an instinct could
possibly
....
have originated; cases, in which no intermediate gradations are known to exist; cases of instinct of
apparently
....
such trifling importance, that they could hardly have been acted on by natural selection; cases of instincts almost identically the same in animals so remote in the scale of nature, that we cannot account for their similarity by inheritance from a common
parent,
progenitor,
and
must therefore
consequently must
believe that they were independently acquired through natural selection. I will not here enter on these several cases, but will confine myself to one special difficulty, which at first appeared to me insuperable, and actually fatal to
my
the
whole theory. I allude to the neuters or sterile females in insect-communities: for these neuters often differ widely in instinct and in structure