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conditions long-continued should likewise induce this 1872
circumstances should both induce and eliminate the same 1869

1 blocks not present in 1861 1869 1872; present in 1859 1860 1866
In the second place, some eminent naturalists believe that a long course of domestication tends to eliminate sterility in the successive generations of hybrids, which were at first only slightly sterile; and if this be so, we surely ought not to expect to find sterility both appearing and disappearing under nearly the same conditions of life.

during long periods of time to more uniform conditions, 1872
to more uniform conditions during long periods of time, 1869

OMIT 1872
and been slowly modified 1869

is 1861 1866 1869 1872
seems to me 1859 1860

it becomes in the highest degree improbable that similar conditions long-continued should likewise induce this tendency; though in certain cases, with species having a peculiar constitution, sterility might occasionally be thus
induced.
caused.
Thus, as I believe, we can understand why with domesticated animals varieties have not been produced which are mutually sterile; and why with plants only a few such cases, immediately to be given, have been observed.
The real difficulty in our present subject is not, as it appears to me, why domestic varieties have not become mutually infertile when crossed, but why this has so generally occurred with natural
varieties
varieties,
as soon as they have been
modified
permanently modified
in a sufficient
and permanent
....
degree to take rank as species. We are far from precisely knowing the cause; nor is this surprising, seeing how profoundly ignorant we are in regard to the normal and abnormal action of the reproductive system. But we can see that species, owing to their struggle for existence with numerous competitors,
must
will
have been exposed during long periods of time to more uniform conditions, than have
been
....
domestic varieties; and this may well make a wide difference in the result. For we know how commonly wild animals and plants, when taken from their natural conditions and subjected to captivity, are rendered sterile; and the reproductive functions of organic beings which have always lived OMIT under natural conditions would probably in like manner be eminently sensitive to the influence of an unnatural cross. Domesticated productions, on the other hand, which, as shown by the mere fact of their domestication, were not originally highly sensitive to changes in their conditions of life, and which can now generally resist with undiminished fertility repeated changes of conditions, might be expected to produce varieties, which would be little liable to have their reproductive powers injuriously affected by the act of crossing with other varieties which had originated in a like manner.
I have as yet spoken as if the varieties of the same species were
almost invariably
invariably
fertile when intercrossed. But it is impossible to resist the evidence of the existence of a certain amount of sterility in the few following cases, which I will briefly abstract. The evidence is at least as good as that from which we believe in the sterility of a multitude of species. The evidence is, also, derived from hostile witnesses, who in all other cases consider fertility and sterility as safe criterions of specific distinction. Gärtner kept during several years a dwarf kind of maize with yellow seeds, and a tall variety with red
seeds,
seeds
growing near each other in his garden; and although these plants have separated sexes, they never naturally crossed. He then fertilised thirteen flowers of the one
with the
kind with
pollen of the