Origin of Species Variorum

Comparison with 1869

by numerous fine, intermediate fossil links, by asking CHAP. IX. GEOLOGICAL RECORD. ourselves whether, for instance, geologists at some future period will be able to prove, that our different breeds of cattle, sheep, horses, and dogs have descended from a single stock or from several aboriginal stocks; or, again, whether certain sea-shells inhabiting the shores of North America, which are ranked by some conchologists as distinct species from their European representatives, and by other conchologists as only varieties, are really varieties or are, as it is called, specifically distinct. This could be effected only by the future geologist discovering in a fossil state numerous intermediate gradations; and such success seems to me improbable in the highest degree.

It has been asserted over and over again, by writers who believe in the immutability of species, that geology has yielded no linking forms. This assertion is entirely erroneous. As Mr. Lubbock has recently remarked, "Every species is a link between other allied forms." We clearly see this if we take a genus having a score of recent and extinct species and destroy four-fifths of them; for in this case no one will doubt that the remainder will stand much more distinct from each other. If the extreme forms in the genus happen to have been thus destroyed, the genus itself in most cases will stand more distinct from other allied genera. The camel and the pig, or the horse and the tapir, are now obviously very distinct forms; but if we add the several fossil quadrupeds which have already been discovered to the families including the camel and pig, these forms become joined by links not extremely wide apart. The chain of linking forms does not, however, in these cases, or in any case, run straight from the one living form to the other, but takes a circuitous sweep through the forms which lived during long past ages. What geological research has not revealed is the former existence of infinitely numerous gradations, as fine as existing varieties, connecting nearly all our existing species with extinct species. And this not having been effected by geology is the most obvious of the many objections which may be urged against my views. ↑

Hence it will be worth while to sum up the foregoing remarks, under an imaginary illustration. The Malay Archipelago is of about the size of Europe from the North Cape to the Mediterranean, and from Britain to Russia; and therefore equals all the geological formations which have been examined with any accuracy, excepting those of the United States of America. I fully agree with Mr. Godwin-Austen, that the present condition of the Malay Archipelago, with its numerous large islands separated by wide and shallow seas, probably represents the former state of Europe, when most of our formations were accumulating. The Malay Archipelago is one of the richest regions of the whole world in organic beings; yet if all the species were to be collected which have ever lived there, how imperfectly would they represent the natural history of the world!

But we have every reason to believe that the terrestrial productions of the archipelago would be preserved in an excessively imperfect manner in the formations which we suppose to be there accumulating. I suspect that not many of the strictly littoral

by numerous, fine, intermediate, fossil links, by asking ... ourselves whether, for instance, geologists at some future period will be able to prove that our different breeds of cattle, sheep, horses, and dogs are descended from a single stock or from several aboriginal stocks; or, again, whether certain sea-shells inhabiting the shores of North America, which are ranked by some conchologists as distinct species from their European representatives, and by other conchologists as only varieties, are really varieties, or are, as it is called, specifically distinct. This could be effected by the future geologist only by his discovering in a fossil state numerous intermediate gradations; and such success is improbable in the highest degree.

It has been asserted over and over again, by writers who believe in the immutability of species, that geology .. yields no linking forms. This assertion, as we shall see in the next chapter, is certainly erroneous. As Sir J. Lubbock has .. remarked, "Every species is a link between other allied forms." If we take a genus having a score of species, recent and extinct, and destroy four-fifths of them, ... no one .. doubts that the remainder will stand much more distinct from each other. If the extreme forms in the genus happen to have been thus destroyed, the genus itself ... will stand more distinct from other allied genera. ↑ What geological research has not revealed, is the former existence of infinitely numerous gradations, as fine as existing varieties, connecting together nearly all existing and extinct species. But this ought not to be expected; yet this has been repeatedly advanced as a most serious objection against my views. ↑

It may be worth while to sum up the foregoing remarks on the causes of the imperfection of the geological record under an imaginary illustration. The Malay Archipelago is .. about the size of Europe from the North Cape to the Mediterranean, and from Britain to Russia; and therefore equals all the geological formations which have been examined with any accuracy, excepting those of the United States of America. I fully agree with Mr. Godwin-Austen, that the present condition of the Malay Archipelago, with its numerous large islands separated by wide and shallow seas, probably represents the former state of Europe, whilst most of our formations were accumulating. The Malay Archipelago is one of the richest regions ... in organic beings; yet if all the species were to be collected which have ever lived there, how imperfectly would they represent the natural history of the world!

But we have every reason to believe that the terrestrial productions of the archipelago would be preserved in an extremely imperfect manner in the formations which we suppose to be there accumulating. Not many of the strictly littoral