Origin of Species Variorum

Comparison with 1860

CHAPTER X.

ON THE GEOLOGICAL SUCCESSION OF ORGANIC BEINGS.

On the slow and successive appearance of new species— On their different rates of change— Species once lost do not reappear— Groups of species follow the same general rules in their appearance and disappearance as do single species— On Extinction— On simultaneous changes in the forms of life throughout the world— On the affinities of extinct species to each other and to living species— On the state of development of ancient forms— On the succession of the same types within the same areas— Summary of preceding and present chapters.

LET us now see whether the several facts and rules relating to the geological succession of organic beings, better accord with the common view of the immutability of species, or with that of their slow and gradual modification, through descent and natural selection.

New species have appeared very slowly, one after another, both on the land and in the waters. Lyell has shown that it is hardly possible to resist the evidence on this head in the case of the several tertiary stages; and every year tends to fill up the blanks between them, and to make the percentage system of lost and new forms more gradual. In some of the most recent beds, though undoubtedly of high antiquity if measured by years, only one or two species are lost forms, and only one or two are new forms, having here appeared for the first time, either locally, or, as far as we know, on the face of the earth. If we may trust the observations of Philippi in Sicily, the successive changes in the marine inhabitants of that island have been many and most gradual. The secondary formations are more broken; but, as Bronn has remarked, neither the appearance nor disappearance of their many now extinct species has been simultaneous in each separate formation.

Species of different genera and classes have not changed at the same rate, or in the same degree. In the oldest tertiary beds a few living shells may still be found in the midst of a multitude of extinct forms. Falconer has given a striking instance of a similar fact, in an existing crocodile associated with many strange and lost mammals and reptiles in the sub-Himalayan deposits. The Silurian Lingula differs but little from the living species of this genus; whereas most of the other Silurian Molluscs and all the Crustaceans have changed greatly. The productions of the land

CHAPTER XI.

ON THE GEOLOGICAL SUCCESSION OF ORGANIC BEINGS.

On the slow and successive appearance of new species— On their different rates of change— Species once lost do not reappear— Groups of species follow the same general rules in their appearance and disappearance as do single species— On extinction— On simultaneous changes in the forms of life throughout the world— On the affinities of extinct species to each other and to living species— On the state of development of ancient forms— On the succession of the same types within the same areas— Summary of preceding and present chapter.

LET us now see whether the several facts and laws relating to the geological succession of organic beings .. accord best with the common view of the immutability of species, or with that of their slow and gradual modification, through variation and natural selection.

New species have appeared very slowly, one after another, both on the land and in the waters. Lyell has shown that it is hardly possible to resist the evidence on this head in the case of the several tertiary stages; and every year tends to fill up the blanks between the stages, and to make the proportion between the lost and existing forms more gradual. In some of the most recent beds, though undoubtedly of high antiquity if measured by years, only one or two species are extinct, and only one or two are new, .. having appeared there for the first time, either locally, or, as far as we know, on the face of the earth. ↑ The secondary formations are more broken; but, as Bronn has remarked, neither the appearance nor disappearance of the many species embedded in each formation has been simultaneous. .. .. .. ...

Species belonging to different genera and classes have not changed at the same rate, or in the same degree. In the older tertiary beds a few living shells may still be found in the midst of a multitude of extinct forms. Falconer has given a striking instance of a similar fact, for an existing crocodile is associated with many .. lost mammals and reptiles in the sub-Himalayan deposits. The Silurian Lingula differs but little from the living species of this genus; whereas most of the other Silurian Molluscs and all the Crustaceans have changed greatly. The productions of the land