Origin of Species Variorum

Comparison with 1869

On the theory of descent, the full meaning of the fact of fossil remains from closely consecutive formations, though ranked as distinct species, being closely related, is obvious. As the accumulation of each formation has often been interrupted, and as long blank intervals have intervened between successive formations, we ought not to expect to find, as I attempted to show in the last chapter, in any one or two formations all the intermediate varieties between the species which appeared at the commencement and close of these periods; but we ought to find after intervals, very long as measured by years, but only moderately long as measured geologically, closely allied forms, or, as they have been called by some authors, representative species; and these we assuredly do find. We find, in short, such evidence of the slow and scarcely sensible mutation of specific forms, as we have a just right to expect to find.

On the State of Development of Ancient compared with Living Forms.

We have seen in the fourth chapter that the degree of differentiation and specialisation of the parts in all organic beings, when arrived at maturity, is the best standard, as yet suggested, of their degree of perfection or highness. We have also seen that, as the specialisation of parts and organs is an advantage to each being, so natural selection will tend to render the organisation of each being more specialised and perfect, and in this sense higher; not but that it may and will leave many creatures with simple and unimproved structures fitted for simple conditions of life, and in some cases will even degrade or simplify the organisation, yet leaving such degraded beings better fitted for their new walks of life. ↑ ↑ In another and more general manner, new species will become superior to their predecessors; for they will have to beat in the struggle for life all the older forms with which they come into close competition. We may therefore conclude that if under a nearly similar climate the eocene inhabitants of the world could be put into competition with the existing inhabitants, the former would be beaten and exterminated by the latter, as would the secondary by the eocene, and the palæozoic by the secondary forms. ↑ So that by this fundamental test of victory in the battle for life, as well as by the standard of the specialisation of organs, modern forms ought on the theory of natural selection to stand higher than ancient forms. Is this the case? A large majority of palæontologists would certainly answer in the affirmative; and I suppose that the answer must be admitted as true, though difficult of full proof. ↑

It is no valid objection to this conclusion or to the general belief that species in the course of time change,

On the theory of descent, the full meaning of the .. fossil remains from closely consecutive formations being closely related, though ranked as distinct species, is obvious. As the accumulation of each formation has often been interrupted, and as long blank intervals have intervened between successive formations, we ought not to expect to find, as I attempted to show in the last chapter, in any one or in any two formations, all the intermediate varieties between the species which appeared at the commencement and close of these periods: but we ought to find after intervals, very long as measured by years, but only moderately long as measured geologically, closely allied forms, or, as they have been called by some authors, representative species; and these assuredly we do find. We find, in short, such evidence of the slow and scarcely sensible mutations of specific forms, as we have the right to expect. .. ..

On the State of Development of Ancient compared with Living Forms .

We have seen in the fourth chapter that the degree of differentiation and specialisation of the parts in .. organic beings, when arrived at maturity, is the best standard, as yet suggested, of their degree of perfection or highness. We have also seen that, as the specialisation of parts .. is an advantage to each being, so natural selection will tend to render the organisation of each being more specialised and perfect, and in this sense higher; not but that it may .. leave many creatures with simple and unimproved structures fitted for simple conditions of life, and in some cases will even degrade or simplify the organisation, yet leaving such degraded beings better fitted for their new walks of life. ↑ ↑ In another and more general manner, new species .. become superior to their predecessors; for they .. have to beat in the struggle for life all the older forms, with which they come into close competition. We may therefore conclude that if under a nearly similar climate the eocene inhabitants of the world could be put into competition with the existing inhabitants, the former would be beaten and exterminated by the latter, as would the secondary by the eocene, and the palæozoic by the secondary forms. So that by this fundamental test of victory in the battle for life, as well as by the standard of the specialisation of organs, modern forms ought, on the theory of natural selection, to stand higher than ancient forms. Is this the case? A large majority of palæontologists would .. answer in the affirmative; and it seems that this answer must be admitted as true, though difficult of proof.

It is no valid objection to this conclusion, ...