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It is a remarkable fact, strongly insisted on by Hooker in regard to America, and by Alph. de Candolle in regard to Australia, that many more identical plants and allied forms have apparently migrated from the north to the south, than in a reversed direction. We see, however, a few southern vegetable forms on the mountains of Borneo and Abyssinia. I suspect that this preponderant migration from north to south is due to the greater extent of land in the north, and to the northern forms having existed in their own homes in greater numbers, and having consequently been advanced through natural selection and competition to a higher stage of perfection or dominating power, than the southern forms. And thus, when they became commingled during the Glacial period, the northern forms were enabled to beat the less powerful southern forms. Just in the same manner as we see at the present day, that very many European productions cover the ground in La Plata, and in a lesser degree in Australia, and have to a certain extent beaten the natives; whereas extremely few southern forms have become naturalised in any part of Europe, though hides, wool, and other objects likely to carry seeds have been largely imported into Europe during the last two or three centuries from La Plata, and during the last thirty or forty years from Australia. Something of the same kind must have occurred on the intertropical mountains: no doubt before the Glacial period they were stocked with endemic Alpine forms; but these have almost everywhere largely yielded to the more dominant forms, generated in the larger areas and more efficient workshops of the north. In many islands the native productions are nearly equalled or even outnumbered by the naturalised; and if the natives have not been actually exterminated, their numbers have been greatly reduced, and this is the first stage towards extinction. A mountain is an island on the land; and the intertropical mountains before the Glacial period must have been completely isolated; and I believe that the productions of these islands on the land yielded to those produced within the larger areas of the north, just in
case. We must, also, bear in mind the occurrence in both hemispheres of former Glacial periods; for these will account, in accordance with the same principles, for the many quite distinct species inhabiting the same widely separated areas, and belonging to genera not now found in the intermediate torrid zones.
It is a remarkable fact strongly insisted on by Hooker in regard to America, and by Alph. de Candolle in regard to Australia, that many more identical or slightly modified species have migrated from the north to the south, than in a reversed direction. We see, however, a few southern .. forms on the mountains of Borneo and Abyssinia. I suspect that this preponderant migration from the north to the south is due to the greater extent of land in the north, and to the northern forms having existed in their own homes in greater numbers, and having consequently been advanced through natural selection and competition to a higher stage of perfection, or dominating power, than the southern forms. And thus, when the two sets became commingled in the equatorial regions, during the alternations of the Glacial periods, the northern forms were the more powerful and were able to hold their places on the mountains, and afterwards to migrate southward with the southern forms; but not so the southern in regard to the northern forms. In the same manner ... at the present day, we see that very many European productions cover the ground in La Plata, New Zealand, and to a lesser degree in Australia, and have ... beaten the natives; whereas extremely few southern forms have become naturalised in any part of the northern hemisphere, though hides, wool, and other objects likely to carry seeds have been largely imported into Europe during the last two or three centuries from La Plata, and during the last forty or fifty years from Australia. The Neilgherrie mountains in India, however, offer a partial exception; for here, as I hear from Dr. Hooker, Australian forms are rapidly sowing themselves and becoming naturalised. Before the last great Glacial period, no doubt the intertropical mountains ... were stocked with endemic Alpine forms; but these have almost everywhere .. yielded to the more dominant forms, generated in the larger areas and more efficient workshops of the north. In many islands the native productions are nearly equalled, or even outnumbered, by those which have become naturalised; and ... this is the first stage towards their extinction. .. Mountains are islands on the land, and their inhabitants have yielded to those produced within the larger areas of the north, just in