Origin of Species Variorum

Comparison with 1860

within each great class, generally change at a slower rate than the higher forms; and consequently the lower forms will have had a better chance of ranging widely and of still retaining the same specific character. This fact, together with the seeds and eggs of many low forms being very minute and better fitted for distant transportation, probably accounts for a law which has long been observed, and which has lately been admirably discussed by Alph. de Candolle in regard to plants, namely, that the lower any group of organisms is, the more widely it is apt to range.

The relations just discussed,— namely, low and slowly-changing organisms ranging more widely than the high,— some of the species of widely-ranging genera themselves ranging widely,— such facts, as alpine, lacustrine, and marsh productions being related (with the exceptions before specified) to those on the surrounding low lands and dry lands, though these stations are so different,— the very close relation of the distinct species which inhabit the islets of the same archipelago,— and especially the striking relation of the inhabitants of each whole archipelago or island to those of the nearest mainland,— are, I think, utterly inexplicable on the ordinary view of the independent creation of each species, but are explicable on the view of colonisation from the nearest and readiest source, together with the subsequent modification and better adaptation of the colonists to their new homes.

Summary of last and present Chapters .—

In these chapters I have endeavoured to show, that if we make due allowance for our ignorance of the full effects of all the changes of climate and of the level of the land, which have certainly occurred within the recent period, and of other similar changes which may have occurred within the same period; if we remember how profoundly ignorant we are with respect to the many and curious means of occasional transport,— a subject which has hardly ever been properly experimentised on; if we bear in mind how often a species may have ranged continuously over a wide area, and then have become extinct in the intermediate tracts, I think the difficulties in believing that all the individuals of the same species, wherever located, have descended from the same parents, are not insuperable. And we are led to this conclusion, which has been arrived at by many naturalists under the designation of single centres of creation, by some general considerations, more especially from the importance of barriers and from the analogical distribution of sub-genera, genera, and families.

With respect to the distinct species of the same genus, which on my theory must have spread from one parent-source; if we make the same allowances as before for our ignorance, and remember that some forms of life change most slowly,

within each great class the lower organisms change at a slower rate than the higher; .. consequently they .. will have had a better chance of ranging widely and of still retaining the same specific character. This fact, together with that of the seeds and eggs of most lowly organised forms being very minute and better fitted for distant transportal, probably accounts for a law which has long been observed, and which has lately been .. discussed by Alph. de Candolle in regard to plants, namely, that the lower any group of organisms stands, the more widely it ... ranges.

The relations just discussed,— namely, lower organisms ranging more widely than the higher,— some of the species of widely-ranging genera themselves ranging widely,— such facts, as alpine, lacustrine, and marsh productions being generally related to those which live on the surrounding low lands and dry lands,— the striking relationship between the inhabitants of islands and those of the nearest mainland— the still closer relationship of the distinct inhabitants of the islands in the same archipelago— are inexplicable on the ordinary view of the ... independent creation of each species, but are explicable if we admit colonisation from the nearest or readiest source, together with the subsequent ... adaptation of the colonists to their new homes.

Summary of the last and present Chapters .

In these chapters I have endeavoured to show, that if we make due allowance for our ignorance of the full effects of .. changes of climate and of the level of the land, which have certainly occurred within the recent period, and of other .. changes which .. have probably occurred,— if we remember how .. ignorant we are with respect to the many .. curious means of occasional transport,— if we bear in mind, and this is a very important consideration, how often a species may have ranged continuously over a wide area, and then have become extinct in the intermediate tracts,— the difficulty is not insuperable in believing that all the individuals of the same species, wherever found, are descended from common parents. And we are led to this conclusion, which has been arrived at by many naturalists under the designation of single centres of creation, by various general considerations, more especially from the importance of barriers of all kinds, and from the analogical distribution of sub-genera, genera, and families.

With respect to .. distinct species belonging to the same genus, which on our theory .. have spread from one parent-source; if we make the same allowances as before for our ignorance, and remember that some forms of life have changed very slowly,