Origin of Species Variorum

Comparison with 1869

existing in scanty numbers— in the several great geographical provinces of the world.

On these same principles, we can understand, as I have endeavoured to show, why oceanic islands should have few inhabitants, but of these a great number should be endemic or peculiar; and why, in relation to the means of migration, one group of beings, even within the same class, should have all its species endemic, and another group should have all its species the same with those in other quarters of the world. We can see why whole groups of organisms, as batrachians and terrestrial mammals, should be absent from oceanic islands, whilst the most isolated islands possess their own peculiar species of aërial mammals or bats. We can see why there should be some relation between the presence, in islands, of mammals, in a more or less modified condition, and the depth of the sea between an island and the mainland. We can clearly see why all the inhabitants of an archipelago, though specifically distinct on the several islets, should be closely related to each other, and likewise be related, but less closely, to those of the nearest continent or other source whence immigrants were probably derived. We can see why in two areas, however distant from each other, where very closely allied or representative species exist, there should almost always exist some identical species. ...

As the late Edward Forbes often insisted, there is a striking parallelism in the laws of life throughout time and space: the laws governing the succession of forms in past times being nearly the same with those governing at the present time the differences in different areas. We see this in many facts. The endurance of each species and group of species is continuous in time; for the exceptions to the rule are so few, that they may fairly be attributed to our not having as yet discovered in an intermediate deposit the forms which are therein absent, but which occur above and below: so in space, it certainly is the general rule that the area inhabited by a single species, or by a group of species, is continuous; and the exceptions, which are not rare, may, as I have attempted to show, be accounted for by migration at some former period under different conditions or by occasional means of transport, and by the species having become extinct in the intermediate tracts. Both in time and space, species and groups of species have their points of maximum development. Groups of species, belonging either to a certain period of time, or to a certain area, are often characterised by trifling characters in common, as of sculpture or colour. In looking to the long succession of ages, as in now

existing in scanty numbers— and this we do find in the several great geographical provinces of the world.

On these same principles we can understand, as I have endeavoured to show, why oceanic islands should have few inhabitants, but that of these, a large proportion should be endemic or peculiar; and why, in relation to the means of migration, one group of beings ... should have all its species peculiar, and another group, even within the same class, should have all its species the same with those in an adjoining quarter of the world. We can see why whole groups of organisms, as batrachians and terrestrial mammals, should be absent from oceanic islands, whilst the most isolated islands should possess their own peculiar species of aerial mammals or bats. We can see why, in islands, there should be some relation between the presence of mammals, in a more or less modified condition, and the depth of the sea between such islands and the mainland. We can clearly see why all the inhabitants of an archipelago, though specifically distinct on the several islets, should be closely related to each other; and should likewise be related, but less closely, to those of the nearest continent, or other source whence immigrants might have been derived. We can see why, if there exist very closely allied or representative species in two areas, however distant from each other, some identical species will almost always there be found.

As the late Edward Forbes often insisted, there is a striking parallelism in the laws of life throughout time and space; the laws governing the succession of forms in past times being nearly the same with those governing at the present time the differences in different areas. We see this in many facts. The endurance of each species and group of species is continuous in time; for the apparent exceptions to the rule are so few, that they may fairly be attributed to our not having as yet discovered in an intermediate deposit certain forms which are .. absent in it, but which occur both above and below: so in space, it certainly is the general rule that the area inhabited by a single species, or by a group of species, is continuous, and the exceptions, which are not rare, may, as I have attempted to show, be accounted for by former migrations under different circumstances, or through occasional means of transport, or by the species having become extinct in the intermediate tracts. Both in time and space, species and groups of species have their points of maximum development. Groups of species, living during the same period of time, or living within the same area, are often characterised by trifling features in common, as of sculpture or colour. In looking to the long succession of past ages, as in ..