Origin of Species Variorum

Comparison with 1861

allied genera, which lived during the Silurian epoch, and these have descended from a species which existed at an unknown anterior period. Species of three of these genera (A, F, and I) have transmitted modified descendants to the present day, represented by the fifteen genera ( a 14 to z 14 ) on the uppermost horizontal line. Now all these modified descendants from a single species, are represented as related in blood or descent to the same degree; they may metaphorically be called cousins to the same millionth degree; yet they differ widely and in different degrees from each other. The forms descended from A, now broken up into two or three families, constitute a distinct order from those descended from I, also broken up into two families. Nor can the existing species, descended from A, be ranked in the same genus with the parent A; or those from I, with the parent I. But the existing genus F 14 may be supposed to have been but slightly modified; and it will then rank with the parent-genus F; just as some few still living organic beings belong to Silurian genera. So that the amount or value of the differences between organic beings all related to each other in the same degree in blood, has come to be widely different. Nevertheless their genealogical arrangement remains strictly true, not only at the present time, but at each successive period of descent. All the modified descendants from A will have inherited something in common from their common parent, as will all the descendants from I; so will it be with each subordinate branch of descendants, at each successive period. If, however, we choose to suppose that any of the descendants of A or of I have been so much modified as to have more or less completely lost traces of their parentage, in this case, their places in a natural classification will have been more or less completely lost,— as sometimes seems to have occurred with existing organisms. All the descendants of the genus F, along its whole line of descent, are supposed to have been but little modified, and they yet form a single genus. ↑ But this genus, though much isolated, will still occupy its proper intermediate position; for F originally was intermediate in character between A and I, and the several genera descended from these two genera will have inherited to a certain extent their characters. This natural arrangement is shown, as far as is possible on paper, in the diagram, but in much too simple a manner. If a branching diagram had not been used, and only the names of the groups had been written in a linear series, it would have been still less possible to have given a natural arrangement; and it is notoriously not possible to represent in a series, on a flat surface, the affinities which we discover in nature amongst the beings of the same group. Thus, on the view which I hold, the natural system is genealogical in its arrangement, like a pedigree; but the degrees of modification which the different groups have undergone, have to be expressed by ranking them under different so-called genera, sub-families, families, sections, orders, and classes.

It may be worth while to illustrate this view of classification, by

allied genera existing during the Silurian epoch, and descended from some still earlier form. In three of these genera (A, F, and I), a species has transmitted modified descendants to the present day, represented by the fifteen genera ( a 14 to z 14 ) on the uppermost horizontal line. Now all these modified descendants from a single species, are .. related in blood or descent in the same degree; they may metaphorically be called cousins to the same millionth degree; yet they differ widely and in different degrees from each other. The forms descended from A, now broken up into two or three families, constitute a distinct order from those descended from I, also broken up into two families. Nor can the existing species, descended from A, be ranked in the same genus with the parent A; or those from I, with the parent I. But the existing genus F 14 may be supposed to have been but slightly modified; and it will then rank with the parent-genus F; just as some few still living organisms belong to Silurian genera. So that the comparative value of the differences between these organic beings, which are all related to each other in the same degree in blood, has come to be widely different. Nevertheless their genealogical arrangement remains strictly true, not only at the present time, but at each successive period of descent. All the modified descendants from A will have inherited something in common from their common parent, as will all the descendants from I; so will it be with each subordinate branch of descendants, at each successive stage. If, however, we suppose any descendant of A, or of I, to have become so much modified as to have lost all traces of its parentage, in this case, its place in the natural system will be lost, as seems to have occurred with some few existing organisms. All the descendants of the genus F, along its whole line of descent, are supposed to have been but little modified, and they .. form a single genus. But this genus, though much isolated, will still occupy its proper intermediate position. The representation of the groups, as here given in the diagram on a flat surface, is much too simple. The branches ought to have diverged in all directions. ↑ If ... the names of the groups had been simply written down in a linear series, the representation would have been still less natural; and it is notoriously not possible to represent in a series, on a flat surface, the affinities which we discover in nature amongst the beings of the same group. Thus, .. the ... natural system is genealogical in its arrangement, like a pedigree: but the amount of modification which the different groups have undergone has to be expressed by ranking them under different so-called genera, sub-families, families, sections, orders, and classes.

It may be worth while to illustrate this view of classification, by