Origin of Species Variorum

do believe that natural selection will generally act very slowly, only at long intervals of time, and only on .. a .. few of the inhabitants of the same region. .. .. .. .. I further believe that these slow, intermittent results accord well with what geology tells us of the rate and manner at which the inhabitants of the world have changed.

Slow though the process of selection may be, if feeble man can do much by ... artificial selection, I can see no limit to the amount of change, to the beauty and .. complexity of the coadaptations between all organic beings, one with another and with their physical conditions of life, which may have been affected in the long course of time through nature's power of selection, that is by the survival of the fittest.

Extinction caused by Natural Selection.

This subject will be more fully discussed in our chapter on Geology; but it must here be alluded to from being intimately connected with natural selection. Natural selection acts solely through the preservation of variations in some way advantageous, which consequently endure. Owing to the high geometrical rate of increase of all organic beings, each area is already fully stocked with inhabitants; and it follows from this, that as the favoured forms increase in number, so, generally, will the less favoured .. decrease and become rare. Rarity, as geology tells us, is the precursor to extinction. We can .. see that any form which is represented by few individuals will run a good chance of utter extinction, during great fluctuations in the nature of the seasons, or from a temporary increase in the number of its enemies. .. .. .. .. .. .. .. But we may go further than this; for, as new forms are ... produced, unless we admit that ... specific forms can go on indefinitely increasing in number, many old forms must become extinct. That the number of specific forms has not indefinitely increased, geology plainly tells us; and we shall presently attempt to show why it is that the number of species throughout the world has not become immeasurably great. ↑

We have seen that the species which are most numerous in individuals have the best chance of producing favourable variations within any given period. We have evidence of this, in the facts stated in the second chapter, showing that it is the common and diffused or dominant species which offer the greatest number of recorded varieties. .. .. .. Hence, rare species will be less quickly modified or improved within any given period; .. they will consequently be beaten in the race for life by the modified and improved descendants of the commoner species.

From these several considerations I think it inevitably follows,