Origin of Species Variorum

Comparison with 1869

care not how trifling a character may be— let it be the mere inflection of the angle of the jaw, the manner in which an insects wing is folded, whether the skin be covered by hair or feathers— if it prevail throughout many and different species, especially those having very different habits of life, it assumes high value; for we can account for its presence in so many forms with such different habits, only by .. inheritance from a common parent. We may err in this respect in regard to single points of structure, but when several characters, let them be ever so trifling, concur .. throughout a large group of beings having different habits, we may feel almost sure, on the theory of descent, that these characters have been inherited from a common ancestor. And we know that such correlated or aggregated characters have especial value in classification.

We can understand why a species or a group of species may depart, in several of its most important characteristics, from its allies, and yet be safely classed with them. This may be safely done, and is often done, as long as a sufficient number of characters, let them be ever so unimportant, betrays the hidden bond of community of descent. Let two forms have not a single character in common, yet if these extreme forms are connected together by a chain of intermediate groups we may at once infer their community of descent, and we put them all into the same class. As we find organs of high physiological importance— those which serve to preserve life under the most diverse conditions of existence— are generally the most constant, we attach especial value to them; but if these same organs, in another group or section of a group, are found to differ much, we at once value them less in our classification. We shall presently see why embryological characters are of such high classificatory importance.

care not how trifling a character may be— let it be the mere inflection of the angle of the jaw, the manner in which an insect's wing is folded, whether the skin be covered by hair or feathers— if it prevail throughout many and different species, especially those having very different habits of life, it assumes high value; for we can account for its presence in so many forms with such different habits, only by its inheritance from a common parent. We may err in this respect in regard to single points of structure, but when several characters, let them be ever so trifling, occur together throughout a large group of beings having different habits, we may feel almost sure, on the theory of descent, that these characters have been inherited from a common ancestor. And we know that such correlated or aggregated characters have especial value in classification.

We can understand why a species or a group of species may depart, in several of its most important characteristics, from its allies, and yet be safely classed with them. This may be safely done, and is often done, as long as a sufficient number of characters, let them be ever so unimportant, betrays the hidden bond of community of descent. Let two forms have not a single character in common, yet if these extreme forms are connected together by a chain of intermediate groups, we may at once infer their community of descent, and we put them all into the same class. As we find organs of high physiological importance— those which serve to preserve life under the most diverse conditions of existence— are generally the most constant, we attach especial value to them; but if these same organs, in another group or section of a group, are found to differ much, we at once value them less in our classification. We shall hereafter, I think, clearly see why embryological characters are of such high classificatory importance.