part
or organ or organ 1859 1860 1861 1866 | or organ 1869 1872 |
is homologous with another
in in 1859 1860 1861 1866 | part in 1869 1872 |
the same individual. And we can understand this fact; for in molluscs, even in the lowest members of the class, we do not find nearly so much indefinite repetition of any one
part, part, 1859 1860 1861 1866 1869 | part 1872 |
as we find in the other great classes of the animal and vegetable kingdoms. ↑6 blocks not present in 1859 1860 1861 1866 1869; present in 1872 | But morphology is a much more complex subject than it at first appears, as has lately been well shown in a remarkable paper by Mr. E. Ray Lankester, who has drawn an important distinction between certain classes of cases which have all been equally ranked by naturalists as homologous.
He proposes to call the structures which resemble each other in distinct animals, owing to their descent from a common progenitor with subsequent modification,
homogenous;
and the resemblances which cannot thus be accounted for, he proposes to call
homoplastic
. For instance, he believes that the hearts of birds and mammals are as a whole homogenous,— that is, have been derived from a common progenitor; but that the four cavities of the heart in the two classes are homoplastic,— that is, have been independently developed.
Mr. Lankester also adduces the close resemblance of the parts on the right and left sides of the body, and in the successive segments of the same individual animal; and here we have parts commonly called homologous, which bear no relation to the descent of distinct species from a common progenitor.
Homoplastic structures are the same with those which I have classed, though in a very imperfect manner, as analogous modifications or resemblances.
Their formation may be attributed in part to distinct organisms, or to distinct parts of the same organism, having varied in an analogous manner; and in part to similar modifications, having been preserved for the same general purpose or function,— of which many instances have been given.
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Naturalists frequently speak of the skull as formed of metamorphosed
vertebræ: vertebræ: 1859 1860 1861 1866 | vertebræ; 1869 1872 |
the jaws of crabs as metamorphosed legs; the stamens and pistils
of of 1859 1860 1861 1866 | in 1869 1872 |
flowers as metamorphosed leaves; but it would in
these these 1859 1860 1861 1866 | most 1869 1872 |
cases
probably probably 1859 1860 1861 1866 1869 | probably 1872 |
be more correct, as Professor Huxley has remarked, to speak of both skull and vertebræ,
both both 1859 1860 1861 1866 1869 | both 1872 |
jaws and legs,
&c.,— &c.,— 1859 1860 1861 | &c., 1866 1869 1872 |
as having been metamorphosed, not one from the
other, other, 1859 1860 1861 1869 1872 | other 1866 |
but but 1859 1860 1861 |
in their present state, but 1866 |
as they now exist, but 1869 1872 |
from some common
element. element. 1859 1860 1861 |
and simpler element. 1866 1869 1872 |
Naturalists, Naturalists, 1859 1860 1861 | Most naturalists, 1866 1869 1872 |
however, use such language only in a metaphorical
sense: sense: 1859 1860 1861 1866 | sense; 1869 1872 |
they are far from meaning that during a long course of descent, primordial organs of any kind— vertebræ in the one case and legs in the other— have actually been
modified modified 1859 1860 1861 1866 | converted 1869 1872 |
into skulls or jaws. Yet so strong is the appearance of
a modification of this nature having a modification of this nature having 1859 1860 1861 |
such modifications having actually 1866 |
this having 1869 1872 |
occurred, that naturalists can hardly avoid employing language having this plain signification.
On my view these terms On my view these terms 1859 1860 1861 1866 |
According to the views here maintained, such language 1869 1872 |
may be used literally; and the wonderful fact of the jaws, for instance, of a crab retaining numerous characters, which they
would probably would probably 1859 1860 1861 | probably would 1866 1869 1872 |
have retained through inheritance, if they had really been metamorphosed
during a long course of descent during a long course of descent 1859 1860 1861 1866 |
OMIT 1869 1872 |
from true
legs, or from some simple appendage, is legs, or from some simple appendage, is 1859 1860 1861 |
though simple legs, is 1866 |
though extremely simple legs, is 1869 |
though extremely simple legs, is in part 1872 |
explained. ↑Subtitle not present 1859 1860 1861 |
Embryology and
Development
.
1866 1869 1872 |
↑1 blocks not present in 1859 1860 1861; present in 1866 1869 1872 | This is one of the most important departments
of natural history.
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Embryology
.—
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It has already been
casually remarked that certain organs casually remarked that certain organs 1859 1860 1861 |
remarked that various parts and organs of the same individual animal are during an early embryonic period exactly like each other, but become 1866 |
stated that various parts and organs 1869 |
stated that various parts 1872 |
in the
individual, which when mature become widely different and serve for different purposes, are in the embryo exactly alike. individual, which when mature become widely different and serve for different purposes, are in the embryo exactly alike. 1859 1860 1861 |
adult state widely different and serve for widely different purposes. 1866 |
same individual are exactly like each other during an early embryonic period, but in the adult state become widely different and serve for widely different purposes. 1869 |
same individual which are exactly alike during an early embryonic period, become widely different and serve for widely different purposes in the adult state. 1872 |
The embryos, also, of distinct animals within the same class are often strikingly similar: a better proof of this cannot be given, than a
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