Origin of Species Variorum

Comparison with 1869

rudiments represent wings. Rudimentary organs sometimes retain their potentiality: this occasionally occurs with the mammæ of male mammals, for they have been known to become well developed, and to secrete milk. So again ... in the udders in the genus Bos, there are normally four developed and two rudimentary teats; but the latter in our domestic cows sometimes become well developed and yield milk. In regard to plants the petals are sometimes rudimental, and sometimes well-developed in individuals of the same species. In certain diœcious plants Kölreuter found that by crossing a species, in which the male flowers included a rudiment of a pistil, with an hermaphrodite species, having of course a well-developed pistil, the rudiment in the hybrid offspring was much increased in size; and this clearly shows that the rudimentary and .. perfect pistils are essentially alike in nature. An animal may possess various parts in a perfect state, and yet they may in one sense be rudimentary, for they are useless: thus the tadpole of the common Salamander or newt, as Mr. G. H. Lewes remarks, "has gills, and passes its existence "in the water; but the Salamandra atra, which lives "high up among the mountains, brings forth its young "full-formed. This animal never lives in the water. "Yet if we open a gravid female, we find tadpoles "inside her with exquisitely feathered gills; and when "placed in water they swim about like the tadpoles of the "water-newt. Obviously this aquatic organisation has "no reference to the future life of the animal, nor has "it any adaptation to its embryonic condition; it has "solely reference to ancestral adaptations, it repeats a "phase in the development of its progenitors."

An organ, serving for two purposes, may become rudimentary or utterly aborted for one, even the more important purpose, and remain perfectly efficient for the other. Thus in plants, the office of the pistil is to allow the pollen-tubes to reach the ovules within the ovarium. .. .. .. The pistil consists of a stigma supported on a style; but in some Compositæ, the male florets, which of course cannot be fecundated, have a rudimentary pistil, for it is not crowned with a stigma; but the style remains well developed, and is clothed with hairs, in the usual manner, for brushing the pollen out of the surrounding and conjoined anthers. Again, an organ may become rudimentary for its proper purpose, and be used for a distinct object: in certain fish the swimbladder seems to be rudimentary for its proper function of giving buoyancy, but has become converted into a nascent

rudiments represent wings. Rudimentary organs sometimes retain their potentiality, and are merely not developed: this seems to be the case with the mammæ of male mammals, for many instances are on record of these organs having become well developed in full-grown males, and having secreted milk. So again there are normally four developed and two rudimentary teats in the udders of the genus Bos, but in our domestic cows the two sometimes become developed and give milk. In plants of the same species the petals sometimes occur as mere rudiments, and sometimes in a well-developed state. In plants with separated sexes, the male flowers often have a rudiment of a pistil; and Kölreuter found that by crossing such male plants with an hermaphrodite species, the rudiment of the pistil in the hybrid offspring was much increased in size; and this shows that the rudiment and the perfect pistil are essentially alike in nature. ↑

An organ serving for two purposes, may become rudimentary or utterly aborted for one, even the more important purpose; and remain perfectly efficient for the other. Thus in plants, the office of the pistil is to allow the pollen-tubes to reach the ovules protected in the ovarium at its base. The pistil consists of a stigma supported on the style; but in some Compositæ, the male florets, which of course cannot be fecundated, have a pistil, which is in a rudimentary state, for it is not crowned with a stigma; but the style remains well developed, and is clothed with hairs as in other compositæ, for the purpose of brushing the pollen out of the surrounding anthers. Again, an organ may become rudimentary for its proper purpose, and be used for a distinct object: in certain fish the swim-bladder seems to be nearly rudimentary for its proper function of giving buoyancy, but has become converted into a nascent