Origin of Species Variorum

Introduction

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the high importance of looking to types, whether or not we can separate and define the groups to which such types belong.

Finally, we have seen that natural selection, which results from the struggle for existence, and which almost inevitably induces extinction and divergence of character in the many descendants from one dominant parent-species, explains that great and universal feature in the affinities of all organic beings, namely, their sub-ordination in group under group. We use the element of descent in classing the individuals of both sexes and of all ages, although having but few characters in common, under one species; we use descent in classing acknowledged varieties, however different they may be from their parent; and I believe this element of descent is the hidden bond of connexion which naturalists have sought under the term of the Natural System. On this idea of the natural system being, in so far as it has been perfected, genealogical in its arrangement, with the grades of difference between the descendants from a common parent, expressed by the terms genera, families, orders, &c., we can understand the rules which we are compelled to follow in our classification. We can understand why we value certain resemblances far more than others; why we are permitted to use rudimentary and useless organs, or others of trifling physiological importance; why, in comparing one group with a distinct group, we summarily reject analogical or adaptive characters, and yet use these same characters within the limits of the same group. We can clearly see how it is that all living and extinct forms can be grouped together in one great system; and how the several members of each class are connected together by the most complex and radiating lines of affinities. We shall never, probably, disentangle the inextricable web of affinities between the members of any one class; but when we have a distinct object in view, and do not look to some unknown plan of creation, we may hope to make sure but slow progress.

Text in this page (from paragraph 2810, sentence 100 to paragraph 2810, sentence 300, word 19) is not present in 1866

the high importance of looking to types, whether or not we can separate and define the groups to which such types belong.

Finally, we have seen that natural selection, which follows from the struggle for existence, and which almost inevitably leads to extinction and divergence of character in the descendants from any one .. parent-species, explains that great and universal feature in the affinities of all organic beings, namely, their subordination in group under group. We use the element of descent in classing the individuals of both sexes and of all ages ... under one species, although they may have but few characters in common; we use descent in classing acknowledged varieties, however different they may be from their parents; and I believe that this element of descent is the hidden bond of connexion which naturalists have sought under the term of the Natural System. On this idea of the natural system being, in so far as it has been perfected, genealogical in its arrangement, with the grades of difference ... expressed by the terms genera, families, orders, &c., we can understand the rules which we are compelled to follow in our classification. We can understand why we value certain resemblances far more than others; why we ... use rudimentary and useless organs, or others of trifling physiological importance; why, in finding the relations between one group and another, we summarily reject analogical or adaptive characters, and yet use these same characters within the limits of the same group. We can clearly see how it is that all living and extinct forms can be grouped together within a few great classes; and how the several members of each class are connected together by the most complex and radiating lines of affinities. We shall never, probably, disentangle the inextricable web of the affinities between the members of any one class; but when we have a distinct object in view, and do not look to some unknown plan of creation, we may hope to make sure but slow progress.

Professor Häckel in his 'Generelle Morphologie' and in .. other works, has recently brought his great knowledge and abilities to bear on what he calls phylogeny, or the lines of descent of all organic beings. In drawing up the several series he trusts chiefly to embryological characters, but receives aid from homologous and rudimentary organs, as well as from the successive periods at which the various forms of life are believed to have first appeared in our geological formations. He has thus boldly made a great beginning, and shows us how classification will in the future be treated.